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FNA | Family List | FNA Vol. 28 | Fontinalaceae

1. Brachelyma Schimper ex Cardot, Mém. Soc. Sci. Nat. Math. Cherbourg. 28: 37, 130. 1892. • [Greek brachys, short, and elyma, veil, alluding to diminutive calyptra].

Plants small to medium-sized, dull. Stems prostrate or pendent; rhizoids from clusters of initials abaxial to leaf insertions, not or weakly branched; axillary hairs 240-360 µm. Leaves distant proximally, crowded distally, oblong-lanceolate to lanceolate, keeled, conduplicate; margins sometimes narrowly reflexed, entire or serrulate proximally, serrulate to serrate at apex; apex obtuse to acute; costa subpercurrent to percurrent; alar cells quadrate or rectangular, slightly enlarged, walls firm; medial laminal cells rhomboidal, linear-rhomboidal, or elongate. Perigonia lateral in leaf axils. Perichaetia with leaves elongate, sheathing setae. Seta 0.7-1.5 mm. Capsule immersed, oblong-cylindric to oval; annulus massive; operculum long-conic or occasionally obliquely rostrate; endostome trellis imperfect. Calyptra cucullate, covering only operculum. Spores 13-18 µm.

Species 1: c, se United States.

Brachelyma is semi-aquatic; as with many semi-aquatic mosses, the plants can be slender to almost robust. Distinctive features include long, narrow, keeled-conduplicate, sometimes obscurely bordered leaves; strong, single costae; firm-walled, non-bulging alar cells; serrate to serrulate distal leaf margins; often rhomboidal distal laminal cells; long, sheathing perichaetial leaves, and short setae with completely immersed capsules. Curiously, the surface cells of the setae are quadrate to oblate; the result of simple cell elongation in these cells would give the Brachelyma setae the same length as the Dichelyma setae, as well as immersed or exserted capsules.

Lower Taxon


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