[For John Herman Sandberg, 1848-1917, Swedish-born American botanist and physician who collected extensively in the Pacific Northwest]

Ihsan A. Al-Shehbaz

Biennials or perennials; (caudex simple or branched, covered with persistent leaf remains); not scapose; pubescent throughout, trichomes shortly stalked or subsessile, cruciform, Y-shaped, or forked. Stems erect or decumbent, unbranched or branched distally. Leaves basal and cauline; petiolate or sessile; basal usually rosulate, petiolate, blade margins entire, dentate, or lyrate-pinnatifid, (apex obtuse to acute); cauline sessile, blade (base attenuate, not auriculate), margins entire, subentire, dentate, or pinnatifid. Racemes (corymbose, several-flowered), considerably elongated in fruit. Fruiting pedicels ascending to subdivaricate, (straight), slender, (terete). Flowers: sepals (erect), oblong; petals white, oblanceolate-spatulate, (longer than sepals, claw obscurely differentiated from blade, apex rounded); stamens slightly tetradynamous; filaments not dilated basally, (slender); anthers ovate or oblong, (apex obtuse); nectar glands confluent, subtending bases of stamens. Fruits subsessile or shortly stipitate (gynophore less than 1 mm), linear, slightly to strongly torulose, subterete to strongly latiseptate; valves each without midvein or with obscure one on proximal 1/2, sparsely to densely pubescent or glabrescent; replum rounded; septum complete; ovules 12-30 per ovary; (style obsolete or distinct); stigma capitate. Seeds uniseriate, plump, not winged, oblong; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons incumbent. x = 7.

Species 2 (2 in the flora): nw North America.

SELECTED REFERENCE Al-Shehbaz, I. A. 2007b. The North American genus Sandbergia (Boechereae, Brassicaceae). Harvard Pap. Bot. 12: 425-427.

Lower Taxa

• Sandbergia perplexa (L. F. Henderson) Al-Shehbaz
• Sandbergia whitedii (Piper) Greene