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4. Gymnocarpium jessoense (Koidzumi) Koidzumi, Acta Phytotax. Geobot. 5: 40. 1936.
Dryopteris jessoensis Koidzumi, Bot. Mag. (Tokyo) 38: 104. 1924; Aspidium dryopteris (Linnaeus) Baumgarten var. longulum H. Christ; Gymnocarpium longulum (H. Christ) Kitagawa; G. robertianum (Hoffman) Newman subsp. longulum (H. Christ) Toyokuni
Subspecies 2 (1 in the flora): North America, Asia.
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