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FNA | Family List | FNA Vol. 4 | Nyctaginaceae | Boerhavia

8. Boerhavia intermedia M. E. Jones, Contr. W. Bot. 10: 41, unnumbered fig. at end of issue. 1902.

Boerhavia erecta Linnaeus var. intermedia (M. E. Jones) Kearney & Peebles

Herbs, annual; taproot tapered, soft or ± woody. Stems usually erect or ascending, occasionally decumbent, moderately or profusely branched primarily distally, 2-6 (-8) dm, minutely puberulent with bent hairs basally, glabrous or minutely puberulent distally. Leaves mostly in basal 1/2 of plant; larger leaves with petiole 7-25 mm, blade broadly ovate or oval to lanceolate, 20-45 × 7-16 mm (distal leaves smaller, sometimes longer, proportionately narrower), adaxial surface usually glabrous, occasionally glandular-puberulent, often minutely punctate, abaxial surface paler than adaxial, glabrous or glabrate, usually punctate with small patches of reddish or brownish cells, base round, obtuse, or truncate, margins entire or slightly sinuate, apex acute, obtuse, or round. Inflorescences terminal, forked ca. 3-6 times ± evenly (or clearly unevenly), diffuse, usually with sticky internodal bands; branches strongly ascending, terminating in umbels or flowers borne singly, occasionally subumbellate (all pedicels not attaching at same point), rarely irregularly compound umbels. Flowers: pedicel 0.5-3.2 mm; bracts at base of perianth quickly deciduous, (1-)2, narrowly lance-acuminate, 0.5-1 mm; perianth whitish to pale pink or purplish, campanulate distal to constriction, 0.7-1.2[-2] mm; stamens 2-3, included or barely exserted. Fruits 1-15 per cluster, straw colored or gray-brown, obconic, broadly low conic, 2-2.8(-3.2) × 0.7-1.3 mm (l/w: 1.7-3.2), apex nearly truncate, glabrous; ribs (4-)5, acute, slightly rugose or undulate near sulci; sulci 0.2-1 times as wide as base of ribs, coarsely transversely rugose, smooth or very faintly papillate. 2n = 52, ca. 54.

Flowering summer-late fall. Sandy or gravelly areas in deserts and arid grasslands, disturbed areas; [0-]100-1700 m; Ariz., Calif., Nev., N.Mex., Tex.; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora).

Boerhavia intermedia is a wide-ranging and variable species of the arid areas of southwestern North America. Ordinarily, the terminal inflorescence is an umbel of at least a few flowers. Plants with few-flowered umbels often have only one flower at some of the terminal inflorescences. Plants with predominantly or entirely one-flowered terminal inflorescences occasionally occur in the eastern part of the range. To the west, and especially on the Coloradan portion of the Sonoran Desert and on the Baja California peninsula, plants with one-flowered terminal inflorescences are more frequent, and even though those have five-ribbed fruits, they often have been identified as B. triquetra. In that region, such plants may have proportionately broader fruits as the ribs become more winglike. Some plants in southern California bear a few fruits with four angles, and in this respect are intermediate to B. triquetra.


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