Navajo bladderpod

Lesquerella navajoensis O’Kane, Madroño 46: 88, figs. 1, 2. 1999

Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, tri-chomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). Stems several from base (crowded), erect, not exceeding leaves. Basal leaves: usually absent. Cauline leaves: (petiole not differentiated from blade); blade linear-oblanceolate, 3-8(-13) mm, margins entire. Racemes (secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). Fruiting pedicels (ascending to divaricate-ascending, straight), 3.5-6 mm. Flowers: sepals (yellow-green), linear to narrowly triangular, 3.7-4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2-6.5 mm, (claw joined at right angle). Fruits (becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3-5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4-8 per ovary; style 1.8-3 mm. Seeds plump or slightly flattened, (strongly mucilaginous).

Flowering May-early Jun. Pinyon-juniper com-munities on nearly barren outcrops of Todilto Limestone; of conservation concern; 2200-2400 m; Ariz., N.Mex.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.