| 1 |
Fruits annual (maturing at end of 1st season); if fruits not available, key as if biennial. |
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(2) |
| + |
Fruits biennial (maturing at end of 2d season) except in Q. agrifolia, Q. emoryi, Q. hypoleucoides, and Q. pumila. |
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(6) |
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| 2 (1) |
Leaf blade abaxially glabrous except for tufts of tomentum in vein axils or at base. |
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(3) |
| + |
Leaf blade abaxially uniformly pubescent or tomentose, sometimes glabrate. |
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(4) |
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| 3 (2) |
Leaf blade distinctly convex, adaxially rugose; California. |
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2 Quercus agrifolia |
| + |
Leaf blade planar, adaxially not rugose; Arizona to w Texas. |
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4 Quercus emoryi |
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| 4 (2) |
Leaf blade less than 3 times as long as wide; nut ovoid or conic, 15–35 mm; California. |
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2 Quercus agrifolia |
| + |
Leaf blade more than 3 times as long as wide; nut globose or oblong, 9–15 mm; Arizona and east. |
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(5) |
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| 5 (4) |
Leaf blade abaxially tawny or white-tomentose; cup covering 1/3 nut or less; Arizona to w Texas. |
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6 Quercus hypoleucoides |
| + |
Leaf blade abaxially glabrate or gray- to brown-pubescent; cup covering 1/3–2/3 nut; Mississippi to North Carolina. |
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16 Quercus pumila |
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| 6 (1) |
Evergreen trees or shrubs; leaf blade adaxially noticeably rugose, margins entire or spinose, strongly revolute, abaxially densely tawny or white-tomentose |
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6 Quercus hypoleucoides |
| + |
Leaf blade 5–11-lobed with more than 10 awns. |
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(77) |
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| 7 |
Petiole less than 10 mm. |
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(8) |
| + |
Petiole more than 10 mm. |
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(44) |
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| 8 (7) |
Leaf blade more than 4.5 times as long as wide. |
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(9) |
| + |
Leaf blade less than 4.5 times as long as wide. |
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(12) |
| |
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| 9 (8) |
Petiole glabrous. |
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14 Quercus phellos |
| + |
Petiole pubescent. |
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(10) |
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| 10 (9) |
Low shrubs. |
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16 Quercus pumila |
| + |
Trees. |
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(11) |
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| 11 (10) |
Leaf blade narrowly lanceolate, abaxially glabrous or with tufts of tomentum at base; sc Arizona. |
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5 Quercus viminea |
| + |
Leaf blade elliptic or ovate, abaxially uniformly pubescent or tomentose; e Texas to North Carolina. |
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11 Quercus incana |
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| 12 (8) |
Leaf margins entire or merely spinose. |
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(13) |
| + |
Leaf margins shallowly or deeply lobed, not merely spinose. |
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(32) |
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| 13 (12) |
Abaxial leaf surface uniformly pubescent or tomentose. |
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(14) |
| + |
Abaxial leaf surface glabrous or sparingly pubescent. |
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(19) |
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| 14 (13) |
Leaf blade convex, adaxially pubescent or tomentose. |
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2 Quercus agrifolia |
| + |
Leaf blade planar, adaxially essentially glabrous. |
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(15) |
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| 15 (14) |
Leaf blade more than 2.5 times as long as wide. 16. Low shrubs; leaf margins revolute. |
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(16) |
| + |
Leaf blade less than 2.5 times as long as wide. |
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(18) |
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| 16 (15) |
Low shrub; leaf margins revolute |
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16 Quercus pumila |
| + |
Trees; leaf margins not revolute. |
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(17) |
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| 17 (16) |
Petioles and twigs pubescent; inner surface of cup uniformly pubescent. |
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11 Quercus incana |
| + |
Petioles and twigs glabrous; inner surface of cup glabrous or with only ring of hairs around nut scar. |
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17 Quercus imbricaria |
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| 18 (15) |
Leaf blade widest distal to middle, apex broadly obtuse to rounded; e Texas to Georgia. |
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23 Quercus arkansana |
| + |
Leaf blade widest at or proximal to middle, apex acute; North Carolina to Maine. |
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25 Quercus ilicifolia |
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| 19 (13) |
Leaf blade distinctly convex, adaxially rugose. |
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(20) |
| + |
Leaf blade planar, not adaxially rugose. |
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(21) |
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| 20 (19) |
Apex of leaf blade blunt to attenuate; nut more than 15 mm; California. |
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2 Quercus agrifolia |
| + |
Apex of leaf blade obtuse or rounded; nut less than 15 mm; Florida. |
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19 Quercus inopina |
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| 21 (19) |
Trees or shrubs evergreen (often tardily deciduous in Q. laurifolia, Q. hemisphaerica, and Q. nigra). |
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(22) |
| + |
Trees deciduous (often tardily deciduous in Q. laurifolia, Q. hemisphaerica, and Q. nigra). |
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(28) |
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| 22 (21) |
Twigs pubescent. |
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(23) |
| + |
Twigs glabrous. |
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(25) |
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| 23 (22) |
Nut 21–44 mm; California. |
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1 Quercus wislizeni |
| + |
Nut less than 20 mm; Arizona and east. |
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(24) |
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| 24 (23) |
Base of leaf blade cuneate to rounded; Mississippi to South Carolina. |
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18 Quercus myrtifolia |
| + |
Base of leaf blade cordate; Arizona to w Texas . |
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4 Quercus emoryi |
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| 25 (22) |
Leaf blade widest near apex. |
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15 Quercus nigra |
| + |
Leaf blade widest at or proximal to middle. |
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(26) |
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| 26 (25) |
Leaf blade rarely 3 times as long as wide; nut 21–44 mm; California. |
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1 Quercus wislizeni |
| + |
Leaf blade commonly more than 3 times as long as wide; nut 8–16 mm; e Texas to Virginia. |
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(27) |
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| 27 (26) |
Leaf blade leathery, base obtuse or rounded, apex acute or acuminate; trees on dry, sandy uplands. |
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13 Quercus hemisphaerica |
| + |
Leaf blade thin, base attenuate or cuneate, apex obtuse or rounded; trees on low, wet flood plains and bottoms. |
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12 Quercus laurifolia |
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| 28 (21) |
Twigs pubescent. |
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(29) |
| + |
Twigs glabrous. |
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(30) |
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| 29 (28) |
Leaf blade widest distal to middle, apex broadly obtuse to rounded; e Texas to Georgia. |
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23 Quercus arkansana |
| + |
Leaf blade widest proximal to middle, apex acute or long- attenuate; w Texas. |
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8 Quercus robusta |
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| 30 (28) |
Leaf blade widest near apex. |
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15 Quercus nigra |
| + |
Leaf blade widest at or proximal to middle. |
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(31) |
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| 31 (30) |
Leaf blade leathery, base obtuse or rounded, apex acute or acuminate; trees on dry, sandy uplands. |
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13 Quercus hemisphaerica |
| + |
Leaf blade thin, base attenuate or cuneate, apex obtuse or rounded; trees on low, wet flood plains and bottoms. |
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12 Quercus laurifolia |
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| 32 (12) |
Terminal buds 6–12 mm. |
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(33) |
| + |
Terminal buds 2–6 mm. |
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(35) |
| |
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| 33 (32) |
Twigs glabrous. |
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15 Quercus nigra |
| + |
Twigs pubescent. |
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(34) |
| |
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| 34 (33) |
Leaf blade widest near middle, base attenuate to acute, blade decurrent; margin of cup involute. |
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22 Quercus laevis |
| + |
Leaf blade widest near apex, base rounded to cordate, blade not decurrent; margin of cup not involute. |
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24 Quercus marilandica |
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| 35 (32) |
Twigs pubescent. |
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(36) |
| + |
Twigs glabrous. |
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(39) |
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| 36 (35) |
Apex of leaf broadly obtuse to rounded; e Texas to Long Island. |
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(37) |
| + |
Apex of leaf acute to attenuate; w Texas. |
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(38) |
| |
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| 37 (36) |
Terminal buds glabrous or only ciliate on scale margins. |
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23 Quercus arkansana |
| + |
Terminal buds uniformly pubescent. |
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24 Quercus marilandica |
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|
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| 38 (36) |
Base of leaf blade subcordate to broadly obtuse, awns usually fewer than 10. |
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8 Quercus robusta |
| + |
Base of leaf blade rounded to cuneate, awns usually more than 10. |
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10 Quercus gravesii |
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| 39 (35) |
Leaf blade widest distal to middle. |
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(40) |
| + |
Leaf blade widest at or proximal to middle. |
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(41) |
| |
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| 40 (39) |
Apex of leaf blade obtuse to rounded or blunt. |
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15 Quercus nigra |
| + |
Apex of leaf blade oblong or acute. |
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27 Quercus georgiana |
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| 41 (39) |
Petiole rarely more than 5mm; margins of leaf blade rarely with more than 4 awns; trees evergreen or tardily deciduous. |
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13 Quercus hemisphaerica |
| + |
Petiole commonly more than 5 mm; margins of leaf blade often with more than 4 awns; trees deciduous. |
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(42) |
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| 42 (41) |
Mature leaf blade abaxially with conspicuous (readily discernible to naked eye) tufts of tomentum in vein axils; bark gray to light brown, scaly; Alabama to South Carolina. |
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27 Quercus georgiana |
| + |
Mature leaf blade abaxially with minute (often detectable only with magnification) tufts of tomentum in vein axils (or such tufts absent); bark brown to black, roughly furrowed; w Texas. |
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(43) |
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| 43 (42) |
Base of leaf blade subcordate to broadly obtuse, awns usually fewer than 10. |
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8 Quercus robusta |
| + |
Base of leaf blade rounded to cuneate, awns usually more than 10. |
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10 Quercus gravesii |
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| 44 (7) |
Leaf margins entire or spinose or with 3–7 lobes separated by shallow (less than 1/3 distance to midrib) sinuses; if lobed, each lobe generally with single awn. |
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(45) |
| + |
Leaf margins with 3–11 lobes separated by deep (more than 1/3 distance to midrib) sinuses; lobes generally with 2 or more awns. |
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(61) |
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| 45 (44) |
Leaf blade abaxially pubescent to tomentose. |
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(46) |
| + |
Leaf blade abaxially glabrous except for tufts of tomentum in vein axils, especially along midrib. |
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(52) |
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| 46 (45) |
Plants evergreen; California or w Texas. |
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(47) |
| + |
Plants deciduous; e Texas to New England. |
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(48) |
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| 47 (46) |
Leaf blade convex; California. |
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2 Quercus agrifolia |
| + |
Leaf blade planar; w Texas. |
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9 Quercus tardifolia |
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| 48 (46) |
Leaf blade 2.5–4 times as long as wide; twigs glabrous. |
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17 Quercus imbricaria |
| + |
Leaf blade rarely as much as 2.5 times as long as wide; twigs pubescent. |
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(49) |
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| 49 (48) |
Terminal buds uniformly pubescent, generally exceeding 5 mm. |
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(50) |
| + |
Terminal buds glabrous or hairy only at apex, rarely as much as 5 mm. |
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(51) |
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| 50 (49) |
Petiole mostly more than 20mm; tips of cup scales appressed.. |
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20 Quercus falcata |
| + |
Petiole rarely as much as 20 mm; tips of cup scales loose, especially at cup margin. |
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24 Quercus marilandica |
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| 51 (49) |
Apex of leaf blade broadly obtuse or rounded, margins entire or with 2–3 rounded lobes; e Texas to Georgia. |
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23 Quercus arkansana |
| + |
Apex of leaf blade acute or obtuse, margins always with 3–4 acute lobes; North Carolina to Maine. |
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25 Quercus ilicifolia |
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| 52 (45) |
Leaf blade commonly more than 3 times as long as wide. |
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7 Quercus graciliformis |
| + |
Leaf blade rarely 3 times as long as wide. |
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(53) |
| |
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| 53 (52) |
Leaf blade convex, adaxially rugose. |
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2 Quercus agrifolia |
| + |
Leaf blade planar, not adaxially rugose. |
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(54) |
| |
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| 54 (53) |
Plants evergreen; California. |
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1 Quercus wislizeni |
| + |
Plants deciduous; w Texas and east. |
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(55) |
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| 55 (54) |
Leaf blade widest distal to middle. |
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(56) |
| + |
Leaf blade widest at or proximal to middle. |
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(58) |
| |
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| 56 (55) |
Twigs glabrous. |
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27 Quercus georgiana |
| + |
Twigs pubescent. |
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(57) |
| |
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| 57 (56) |
Terminal buds uniformly pubescent. |
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24 Quercus marilandica |
| + |
Terminal buds glabrous or at most ciliate on scale margins. |
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23 Quercus arkansana |
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| 58 (55) |
Leaf blade usually with more than 10 awns. |
|
(59) |
| + |
Leaf blade usually with fewer than 10 awns. |
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(60) |
| |
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| 59 (58) |
Petiole less than 25 mm; bark black and furrowed; cup less than 15mm wide; w Texas. |
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10 Quercus gravesii |
| + |
Petiole more than 25mm; bark gray or dark gray with wide shiny ridges separated by shallow fissures; cup more than 15 mm wide; Oklahoma to Nova Scotia. |
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32 Quercus rubra |
| |
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|
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| 60 (58) |
Base of leaf blade subcordate to broadly obtuse; cup deeply cup-shaped, 6–9 mm high; w Texas. |
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8 Quercus robusta |
| + |
Base of leaf blade cuneate to obtuse; cup shallowly saucer- shaped, 4–6mm high; Alabama to South Carolina. |
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27 Quercus georgiana |
| |
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|
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| 61 (44) |
Leaf blade abaxially uniformly pubescent or tomentose. |
|
(62) |
| + |
Leaf blade abaxially glabrous or with a few scattered hairs, not uniformly pubescent. |
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(65) |
| |
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|
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| 62 (61) |
Nut oblong to broadly ellipsoid, 21–34 mm; tips of cup scales loose; California to Oregon. |
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3 Quercus kelloggii |
| + |
Nut ovoid or subglobose, 9–16 mm; tips of cup scales appressed; Texas and east. |
|
(63) |
| |
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| 63 (62) |
Base of leaf blade rounded or U-shaped, terminal lobe much longer than lateral lobes. |
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20 Quercus falcata |
| + |
Base of leaf blade cuneate to truncate, terminal lobe rarely exceeding lateral lobes. |
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(64) |
| |
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| 64 (63) |
Trees; terminal buds 4–9 mm, light reddish brown, uniformly puberulent |
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21 Quercus pagoda |
| + |
Shrubs; terminal buds 2–4.5 mm, dark reddish brown, puberulent only at apex. |
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25 Quercus ilicifolia |
| |
|
|
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| 65 (61) |
Terminal buds uniformly pubescent. |
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(66) |
| + |
Terminal buds glabrous or pubescent only on distal 1/2. |
|
(67) |
| |
|
|
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| 66 (65) |
Leaf base attenuate or acute, blade decurrent; margin of cup involute. |
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22 Quercus laevis |
| + |
Leaf base obtuse to truncate, blade not decurrent; margin of cup not involute. |
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35 Quercus velutina |
| |
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| 67 (65) |
Terminal buds silvery or reddish pubescent on distal 1/2. |
|
(68) |
| + |
Terminal buds glabrous or with only a few reddish hairs at apex. |
|
(71) |
| |
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|
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| 68 (67) |
Base of leaf blade attenuate or acute, decurrent; petiole commonly less than 20 mm; margin of cup involute. |
|
22 Quercus laevis |
| + |
Base of leaf blade obtuse to truncate, not decurrent; petiole commonly more than 20 mm; margin of cup not involute. |
|
(69) |
| |
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|
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| 69 (68) |
Leaf blade adaxially dull green; sinuses usually extending less than 1/2 distance to midrib, lobes acute to oblong. |
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32 Quercus rubra |
| + |
Leaf blade adaxially glossy green; sinuses usually extending more than 1/2 distance to midrib, lobes distally expanded. |
|
(70) |
| |
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| 70 (69) |
Cup turbinate to hemispheric, scales with broad glossy base, scale margins often strongly concave; nut ovoid to subglobose, with 1 or more concentric rings of pits at apex. |
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33 Quercus coccinea |
| + |
Cup deeply cup-shaped to turbinate, scales pubescent with straight or slightly concave margins; nut ellipsoid to ovoid, without rings of pits at apex. |
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34 Quercus ellipsoidalis |
| |
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|
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| 71 (67) |
Leaf blades mostly wider than long. |
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31 Quercus acerifolia |
| + |
Leaf blades mostly longer than wide. |
|
(72) |
| |
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| 72 (71) |
Twigs and/or terminal buds yellowish gray, gray, or grayish brown. |
|
(73) |
| + |
Twigs and terminal buds brown to dark reddish brown or red. |
|
(75) |
| |
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|
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| 73 (72) |
Mature leaf blade abaxially glabrous or with minute (often detectable only with magnification) tufts of tomentum in vein axils. |
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29 Quercus buckleyi |
| + |
Mature leaf blade abaxially with conspicuous (readily discernible to naked eye) tufts of tomentum in vein axils. |
|
(74) |
| |
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| 74 (73) |
Cup with thin (less than 1.5mm in cross section) walls, deeply goblet-shaped, covering 1/3–1/2 nut, inner surface pubescent. |
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28 Quercus texana |
| + |
Cup with thick (more than 1.5mm in cross section) walls, saucer- or cup-shaped, covering 1/4–1/3 nut, inner surface glabrous or with ring of hairs around scar. |
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30 Quercus shumardii |
| |
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| 75 (72) |
Leaf blade abaxially with conspicuous (readily discernible to naked eye) tufts of tomentum in vein axils. |
|
(76) |
| + |
Leaf blade abaxially glabrous or with minute (often detectable only with magnification) tufts of tomentum in vein axils. |
|
(78) |
| |
|
|
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| 76 (75) |
Leaf blade 3–5-lobed with fewer than 10 awns. |
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27 Quercus georgiana |
| + |
desc needed |
|
|
| |
|
|
|
| 77 (6) |
Cup deeply goblet- shaped, covering 1/3– 1/2 nut; nut broadly ovoid to broadly ellipsoid, rarely less than 16mm. |
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28 Quercus texana |
| + |
Cup shallowly saucer- shaped, covering less than 1/3 nut; nut globose or ovoid, rarely more than 16mm. |
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26 Quercus palustris |
| |
|
|
|
| 78 (75) |
Cup generally more than 12 mm high; cup scales more than 4mm, attenuate or acuminate to acute with loose tips, especially at cup margin; California to Oregon. |
|
3 Quercus kelloggii |
| + |
Cup rarely as much as 12 mm high; cup scales less than 4mm, acute to obtuse with tightly appressed tips; wTexas and east. |
|
(79) |
| |
|
|
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| 79 (78) |
Leaf blade adaxially dull green, abaxially pale green or glaucous; cup 18–30 mm wide; Oklahoma to Nova Scotia. |
|
32 Quercus rubra |
| + |
Leaf blade adaxially shiny or glossy, abaxially light green or coppery green; cup 7–18 mm wide; Texas to Oklahoma. |
|
(80) |
| |
|
|
|
| 80 (79) |
Petiole more than 20 mm; leaf lobes usually distally expanded; c Texas to Oklahoma. |
|
29 Quercus buckleyi |
| + |
Petiole usually less than 20 mm; leaf lobes acute; w Texas. |
|
10 Quercus gravesii |