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1c. Thelypteris Subg. Lastrea (Hooker) Alston, J. Wash. Acad. Sci. 48: 232. 1958.
Nephrodium Marthe ex Michaux subg. Lastrea Hooker, Sp. Fil. 4: 5, 84. 1862
Distinguished from other subgenera by the combination of seasonal leaves, short-creeping to suberect stems hidden by old petiole bases, persistently scaly petioles, pinnate-pinnatifid blades with greatly reduced proximal pinnae, and veins meeting margin above the sinuses. x = 34.
Thelypteris quelpaertensis , T . limbosperma (Allioni) H. P. Fuchs from Europe, and T . elwesii (Baker) Ching from Asia constitute the genus Oreopteris Holub (R. E. Holttum 1981).
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