9a. Codriophorus P. Beauvois sect. Codriophorus
Racomitrium sect. Papillosa (Kindberg) Noguchi; Trichostomum sect. Stenotrichum Chevallier
Stem sparingly or freely irregularly, dichotomously or fasciculately branched, sometimes filiform and almost unbranched, absent short, lateral tuft-like branchlets. Leaves lingulate, elliptic, broadly ovate, ovate- or oblong-lanceolate to broadly or occasionally narrowly lanceolate; margins 1-2(-4)-stratose, recurved, reflexed, incurved or plane on both sides; apices narrowly to broadly rounded-obtuse or long acuminate to acute, muticous or very rarely shortly hyaline-tipped, entire or sharply to bluntly, remotely irregularly toothed, erose-dentate, sinuate to cristate in the upper 1/5-1/4; costa strong and broad, ceasing in mid leaf to subpercurrent, in transverse-section 2-4-stratose in the distal and medial parts, weakly to strongly convex, reniform, crescent-shaped to semi-terete abaxially, flat or convex on the adaxial side, in the basal part 3-6(-7)-layered, situated at the bottom of a shallow or deeper, usually wide-angled groove, lunate, reniform or rectangular and distinctly flattened, strongly convex on the abaxial side; basal marginal laminal cells not differentiated; distal laminal cells 1- to variously 2-stratose, irregularly rounded-quadrate, subquadrate, oval to rounded-hexagonal, strongly papillose to smooth, straight-walled in the distal part.
Species 9 (6 in the flora): North America, South America (Colombia), Eurasia, n Africa, Atlantic Islands (Azores, Canary Islands, Iceland, Madeira).
Members of sect. Codriophorus are recognized by their lingulate, broadly ovate to ovate- or oblong-lanceolate leaves, isodiametric and straight-walled distal laminal cells, and costa that is broad and robust, usually over 100 µm wide at the base. The costa is situated in a shallow, wide-angled groove and is mostly 2-4-stratose in the distal and medial parts, 3-6(-7)-stratose in the basal part, and anatomically strongly differentiated.
