菊科

PENG, Ching-I & CHUNG, Kuo-Fang; LI, Hui-Lin

Artemisia by LING, Yeou-Ruenn & PENG, Ching-I
Astereae by SOEJIMA, Akiko & PENG, Ching-I
Blumea by LEU, Wen-Pen & PENG, Ching-I
Eupatorium by CHUNG, Shih-Wen & PENG, Ching-I
Senecioneae by PENG, Ching-I & CHUNG, Shih-Wen

Hemistepta lyrata Credit: HAST

Herbs or shrubs, rarely trees, sap sometimes milky. Leaves alternate or opposite, rarely whorled, simple or pinnately lobed or divided, estipulate. Flowers 1 to many, arranged in heads, heads solitary or variously arranged into synflorescences. Receptacle variously shaped, either glabrous, setose, or paleaceous, subtended by an involucre of one or more series of bracts (phyllaries). Florets bisexual or unisexual, occasionally neutral. Calyx represented by a pappus of diverse forms or seemingly absent. Corolla gamopetalous, tubular or discoid with 4 or 5 equal teeth or lobes or 2-5 lobes fused and elongate and forming a 2-5 toothed limb or ligule, or corolla bilabiate. Stamens 5, epipetalous, filaments distinct, anthers introrse, connate and forming a tube around style, apex usually appendaged, base often caudate or sagittate. Pistil 1, ovary inferior, 1-celled; style simple, slender, usually bifid; ovule solitary, anatropous. Fruit an achene, often crowned with a setose, paleaceous, or plumose pappus. Endosperm absent.

One of the largest families of flowering plants comprising about 1,535 genera and about 23,000 species, cosmopolitan. About 84 genera and 221 species, plus 20 additional infraspecific taxa in Taiwan.

• Bremer, K. 1994. Asteraceae: Cladistics and Classification. Timber Press, Portland, 752 pp. Cronquist, S. 1955. Phylogeny and taxonomy of the Compositae. Amer. Midl. Naturalist. 53: 478-511.

• Cronquist, S. 1980. Asteraceae Dumort. In L. S. Radford (ed.), Vascular Flora of the Southern United States. University of North Carolina Press, Chapel Hill, Vol. 1: 3-261.

• Hoffmann, O. 1894. Compositae. In A. Engler and K. A. E. Prantl, Die natürlichen Pflanzenfamilien. IV. 5: 87-391.

• Iwatsuki, K., T. Yamazaki, D. E. Boufford & H. Ohba (eds). 1995. Flora of Japan. Vol. 3(b)-Angiospermae, Dicotyledoneae, Sympetalae(b). Kodansha Ltd., Tokyo, 181 pp.

• Kitamura, S. 1937. Compositae Japonicae. Pars prima. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 13: 1-421, pls. 1-35.

• Kitamura, S. 1940. Compositae Japonicae. Pars secunda. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 15(3): 285-446, pls. 1-11.

  Kitamura, S. 1942. Compositae Japonicae. Pars tertia. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 16(3): 155-292, pls. 1-8.

• Kitamura, S. 1955. Compositae Japonicae. Pars quarta. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 22(1): 77-126, pls. 2-4.

• Kitamura, S. 1956. Compositae Japonicae. Pars quinta. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 23(1): 105-168.

• Kitamura, S. 1957. Compositae Japonicae. Pars sexta. Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 24(1): 1-79, pls. 1-7.

• Ling, Y. & C. Shih (eds.). 1983. Fl. Reipubl. Popul. Sin. Vol. 76(1). Compositae (3)-Anthemideae. Science press, Beijing, 149 pp.

• Ling, Y. & C. Shih (eds.). 1983. Fl. Reipubl. Popul. Sin. Vol. 80(1). Compositae (10)-Lactuceae. Science press, Beijing, 342 pp.

• Ling, Y. & C. Shih (eds.). 1987. Fl. Reipubl. Popul. Sin. Vol. 78(1). Compositae (7)-Echinopsideae, Cynareae. Science press, Beijing, 226 pp.

• Ling, Y. & S. W. Liu (eds.). 1989. Fl. Reipubl. Popul. Sin. Vol. 77(2). Compositae (4)-Senecioninae. Science press, Beijing, 188 pp.

• Ling, Y. & Y. L. Chen (eds.). 1985. Fl. Reipubl. Popul. Sin. Vol. 74. Compositae (1)-Vernonieae-Astereae. Science press, Beijing, 388 pp.

• Ling, Y. & Y. R. Ling (eds.). 1991. Fl. Reipubl. Popul. Sin. Vol. 76(2). Compositae (3)-Anthemideae (2). Science press, Beijing, 321 pp.

• Ling, Y. (ed.). 1979. Fl. Reipubl. Popul. Sin. Vol. 75. Compositae (2)-Inuleae-Helenieae. Science press, Beijing, 422 pp.

• Peng, C.-I & C. C. Hsu. 1978. Chromosome numbers in Taiwan Compositae. Bot. Bull. Acad. Sin. 19: 53-66.

• Tseng, Y. C. (ed.). 1996. Fl. Reipubl. Popul. Sin. Vol. 79. Compositae (9)-Mutisieae. Science press, Beijing, 113 pp.
  
  

  Tribe 8. Senecioneae
  1 Leaves chiefly radical, cordate-reniform, long petiolate; stem scapiform with few bract-like leaves.   (2) + Leaves chiefly cauline or radical and cauline, variously shaped.   (4)         2 (1) Plants partially dioecious; heads discoid.   Petasites + Plants not dioecious; heads radiate.   (3)         3 (2) Leaf margins entire or angularly sinuate dentate, involute; achenes hairy.   Farfugium + Leaf margins dentate to serrate, revolute; achenes glabrous.   Ligularia         4 (1) Heads discoid, corolla various in color.   (5) + Heads radiate, corolla yellow.   (11)         5 (4) Leaves on lower stem peltate and palmatifid.   Syneilesis + Leaves not as above.   (6)         6 (5) Leaves cordate-reniform or hastate, long petiolate.   Parasenecio + Leaves not as above.   (7)         7 (6) Involucre without bracteoles at base.   Emilia + Involucre with bracteoles at base.   (8)         8 (7) Heads heterogamous, outer florets pistillate, central florets bisexual.   Erechtites + Heads homogamous, florets all bisexual.   (9)         9 (8) Heads nodding; corolla gradually widened into limb.   Crassocephalum + Heads erect, corolla rather abruptly broadened into limb.   (10)         10 (9) Style branches very long, much exserted beyond corolla; involucre more than 10 mm long.   Gynura + Style branches short, scarcely exserted beyond corolla; involucre 6-7 mm long.   Senecio (S. vulgaris)         11 (4) Involucre with bracteoles at base.   Senecio + Involucre without bracteoles at base.   (11)         12 Leaves pinnatisect.   Nemosenecio + Leaves subentire.   Tephroseris
  List of Keys KEY TO TRIBES Tribe 1. Vernonieae Tribe 2. Eupatorieae Tribe 3. Astereae Tribe 4. Inuleae Tribe 5. Heliantheae Tribe 7. Anthemideae Tribe 8. Senecioneae Tribe 9. Cardueae Tribe 10. Mutisieae Tribe 11. Lactuceae

• List of lower taxa