Begonia kotoensis Hayata
Plant epipetric or sometimes terrestrial; evergreen; acaulenscent with creeping rhizomes, monoecious.
Rhizomes to ___ cm long, to 19 mm across, glabrous.
Stipules persistent, ovate to triangular, apex mucronate or acuminate, margins entire and often reflex, midrib raised ca. 2 mm abaxially, to 21 mm long, 18 mm wide, progressively hirsute toward the apex.
Leaves many, all basal, thick, asymmetric, very rarely peltate, broadly ovate to orbicular, undivided, apex mucronate to acuminate, base cordate, margins irregularly denticulate or serrulate, 7-26 cm long, 6-21 cm wide, glabrous adaxially, remotely scaberulous abaxially; venation palmate; petioles, 7-31 mm long, 10 mm across, glabrous or nearly so.
Inflorescences arise from rhizomes, 17-53 cm long, glabrous; peduncles (inflorescence axis) to 41 cm long, 9 mm across, erect.
Bracts caducous, orbicular to oblate, apex retuse to apiculate, margins entire and often reflex, to 14 mm long, 18 mm wide, glabrous.
Flowers of both sexes white to pinkish, glabrous.
Staminate flowers: tepals 4, outer two obovate to orbicular, 9-25 mm long, 8-22 mm wide, inner two oblanceolate to narrowly obovate, 8-23 mm long, 4-11 mm wide; androecium zygomorphic; stemans 39-58, (shape), yellow, filaments free, 1.2-2.3 mm long; anthers broadly oblong, 1.4-1.8 mm long, 0.9-1.3 mm across.
Pistillate flowers: tepals 5(-6), oblanceolate to broadly obovate, unequal, the largest 8-21 mm long, 7-18 mm wide, the smallest 7-18 mm long, 4-8 mm wide; ovary white to tinged pinksih when blooming, inferior, glaborus; styles 3, yellow, free, 2-lobed / forked once, 4.3-4.8 mm long; stigmas, in a band and spiraled.
Fruits capsular, dehiscence, nodding, trigonous, subequally 3-winged, abaxial wing transversely elliptic, 6-10 mm long, 9-13 mm wide, lateral wings 3-8 mm long, 8-12 mm wide; locules 3, axile placentation, placental branches 2 per locule.
PINGTUNG: Wanteli Rd.-Nanjenshan, Yang et al. 4330 (THAI).
TAITUNG: Lanyu, Mori 2472 (TAIF), Hosokawa8049 (TAI), Ou et al. 9334 (TCF), Chang 8600, 9830, 11491, 11591, 11592, 11868, 17152 (PPI), Ou 2319, 7923 (TCF), Peng 5235, 6618, 7577 (HAST), Peng & Chen 10634, 10779, 10794 (HAST), Koto et al. s. n. 1982 (PPI), Nakahara 1022 (Syntype of B. kotoensis Hay., TAIF), Tanaka s. n. 1931 (NTUF), Jeng s. n. 1977 (TAI), Iwami 460 (TAI), Hsu 4833, 4951 (TAI), Liu et al. 131 (TAI), Yamamoto s. n. 1947 (TAI), Huang 10355 (TAI), Chuang & Hsu s. n. 1985 (TAI), Huang & Kao 6321 (TAI), Huang et al. 9291 (TAI), Yang 3112 (TAI), Sasaki s. n. Jun. 15, s. n. Jul. 1912, s. n. Feb. 7, 1920 (TAI), Liu 2015 (TAI), Huang et al. 10556 (TAI), Huang 2711 (TAI), Jong 924 (TAI); Lutao, Kawakami & Kobuyasi 4823 (TAIF), Chang 16529 (PPI), Kudo & Mori s. n. 1930 (TAI), Huang & Koa 5095, 5321 (TAI), Iwami s. n. 1987.
Stolons: absent
Stipules: persistence
Stipules: shape: ovate, triangular
Stipules: apex: acuminate, mucronate
Stipules: margin: entire
Stipules: length (mm): 21 mm
Stipules: width (mm): 18 mm
Stipules: indumentum: progressively hirsute toward the apex
Leaves: shape: broadly ovate, orbicular
Leaves: apex: acuminate
Leaves: margin: denticulate, serrulate
Leaves: length (cm): 7-26 cm
Leaves: width (cm): 6-21 cm
Leaves: indumentum: glabrous, scaberulous
Venation: palmate
Veins: number: 7-10
Petioles: length (cm): 7-31 cm
Petioles: diameter (mm): 10 mm
Petioles: indumentum: glabrous
Bracts: caducous
Bracts: shape: oblate, orbicular
Bracts: apex: retuse, apiculate
Bracts: margin: entire
Bracts: length (mm): 14 mm
Bracts: width (mm): 18 mm
Bracts: indumentum: glabrous
Inflorescence: length (cm): 17-53 cm
Peduncles: length (cm): 15-41 cm
Peduncles: diameter (mm): 9 mm
Flowers: color: white, pinkish
Male flower petals: number: four
Outer 2 tepals of male flowers: shape: orbicular, obovate
Outer 2 tepals of male flowers: length (mm): 9-25 mm
Outer 2 tepals of male flowers: width (mm): 8-22 mm
Inner 2 tepals of male flowers: shape: oblanceolate, narrowly obovate
Inner 2 tepals of male flowers: length (mm): 8-23 mm
Inner 2 tepals of male flowers: width (mm): 4-11 mm
Stamens: number: 39-58
Stamens: shape: golf-club shaped
Anthers: shape: broadly oblong
Anthers: length (mm): 1.4-1.8 mm
Anthers: diameter (mm): 0.9-1.3 mm
Filaments: longer than anthers, equal to anthers
Filaments: length (mm): 1.2-2.3 mm
Female flower tepals: number: five, six
Female flower tepals: equality of size: unequal
The largest tepals of female flowers: shape:: broadly obovate
The largest tepals of female flowers: length (mm): 8-21 mm
The largest tepals of female flowers: width (mm): 7-18 mm
The smallest tepals of female flowers: shape: oblanceolate
The smallest tepals of female flowers: length (mm): 7-18 mm
The smallest tepals of female flowers: width (mm): 4-8 mm
Styles: number: two
Styles: length (mm): 4.3-4.8 mm
Locules number: three
Fruits: typ: capsule
Fruits: wings: 3-winged
Abaxil wing of fruits: shape: transversely elliptic
Abaxil wing of fruits: length (mm): 6-10 mm
Abaxil wing of fruits: width (mm): 9-13 mm
Lateral wings of fruits: length (mm): 3-8 mm
Lateral wings of fruits: width (mm): 8-12 mm
Seeds: shape: broadly ellipsoid
Seeds: length (mm): 0.33-0.37 mm
Seeds: diameter (mm): 0.22-0.25 mm
Seed apex: shape: rotund
Seed micropylar end: constricted
Chrosome numer 2n=: 60
Elevation (m): 0-450 m
Distribution: Taitung
Plant: growth form: epipetric
Phenology: evergreen
Stems: habit: rhizomatous
