Begonia hayatae Gagnep.
Plant terrestrial; evergreen; with very short rhizomes from which upright stems arise, monoecious. Stems green to reddish brown, to 110 cm long, 19 mm across, glabrous. Rhizomes greenish or reddish, to ___ cm long, ___ mm across, glabrous. Stipules caducous, narrowly deltoid to lanceolate, apex acuminate, margins entire, to 24 mm long, 19 mm wide, glabrous. Leaves many, upper surface green, lower surface greenish or reddish, asymmetric, lanceolate to broadly lanceolate, undivided, apex acuminate to cuspidate, base obtuse, margins irregularly denticulate or serrulate, 9-24 cm long, 3.5-8 cm wide, sparsely hispidous; venation palmate-pinnate; petioles, 3-12 mm long, to 5 mm across, glabrous. Inflorescences arise from stem, glabrous; peduncles (inflorescence axis) strongly reduced, ca. 2 cm long, 1.6 mm across, pendent; pedicel pendent. Bracts caducous, apex mucronate to acuminate, margins entire, to 19 mm long, 12 mm wide, glabrous. Staminate flowers: tepals 4, white, crateriformis, apex obtuse, glabrous, outer two broadly ovate to subglobose, 8-13 mm long, 6-12 mm wide, inner two elliptic, 7-11 mm long, 4-6 mm wide; androecium actinomorphic; filaments free, nearly equal in length, 0.8-1.4 mm long; anthers (number), yellowish, clavate, longer than filaments, 1.7-2.4 mm long, 0.4-0.6 mm across. Pistillate flowers: tepals 6, white, nearly equal, broadly elliptic to rotund, 7-12 mm long, 5-9 mm wide, glabrous; ovary green when blooming, inferior; styles 3, yellowish, free, forked once, yellowish, 4.9-5.4 mm long;; stigmas in a band and spiraled. Fruits berry-like, indehiscence, pendent, green, broadly ellipsoid, ___ mm long, 17-18 mm across, glabrous; indistinctly 3-ridged to conspicuously 3-horned; shallowly 3-grooved between locules; beaks 0-3 mm long. locules 3, axile placentation, placental branches 2 per locule.
TAIPEI: Shihting-Huangtitien, Hsu & Lin 41 (HAST); Huangtitien, Yang 43 (TAI); Hsiaoyii, Chen 108 (HAST); Wulai, Peng 4688, 8355, 1000 (HAST), Sasaki s.n. Aug. 28, 1927 (TAIF), Jeng 2548 (TAI), Sasaki s.n. May 25, 1932 (TAI), Mutuda s.n. Jul. 25, 1918 (TAIF), Lai s.n. Mar. 31, 1971 (NTUF), Chuang 2310 (TAI); Fushan, Nakamuna 638 (TAI), Huang 1277 (TAI); Tatungshan, Kuo 5581 (TAI); Leikungpo-Hsiaoketou, Peng et al. 11965 (HAST), Peng & Hu 12271 (HAST); Kankou, Kao 7904(TAI), Sasaki s.n. Nov. 1, 1932 (TAI); Chengpai-Peichatienshan, Peng 8611 (HAST).
ILAN: Taipingshan, Suzuki s.n., Aug. 15, 1928 (TAI).
TAOYUAN: Peichatienshan, Peng 10132 (HAST), Mori s.n. May 25, 1908 (TAIF); Chiaopanshan, Hayata s.n. Jan. 1912 (TAIF).
HSINCHU: Wuchihshan, Kawakami 1296 (Syntype of B. aptera Hay., TAIF).
MIAOLI: Tahu, Kawakami et al. 57 (TAIF); Sanyii, Susuki s.n. Jul. 28, 1940 (TAI).
TAICHUNG: Tungshih Forest Station, Chen 393 (HAST); Pahsiehshan, Sasaki s.n. Nov. 23, 1922 (TAIF); Lineng, Sasaki s.n. Nov. 21, 1922 (TAI); Taichung, Suzuki s.n. Jul. 17, 1936 (TAIF).
NANTOU: Shuili, Peng & Chen 11952; Huisun Forest Station, Peng 12232 (HAST), Ou 8511 (TCF), Ying 1220 (NTUF); Meiyuan, Sasaki s.n. Nov. 29, 1922 (TAIF); Touchuang, Shimada 14181 (TAIF); Guandaushi, Liu s.n. Sep. 18, 1969 (TCF); Sun-Moon Lake, Ying 4991 (NTUF); Citou, Liu s.n. Apr. 1, 1971 (NTUF).
HUALIEN: Fuyuan Butterfly Valley, Peng & Chen 11653 (HAST); Tatung-Tali, Chen 161 (HAST); Lienhuachih-Taomikengka, Mori s.n. Jul. 6, 1936 (TAI).
Tebbitt, M.C.2003. Taxonomy of Begonia longifolia Blume (Begoniaceae) and related species. Brittonia 55(1): 19–29.
Stolons: absent
Stipules: caducous
Stipules: shape: triangular, narrowly triangular
Stipules: apex: acuminate
Stipules: margin: entire
Stipules: length (mm): 24 mm
Stipules: width (mm): 19 mm
Stipules: indumentum: glabrous
Leaves: shape: lanceolate
Leaves: apex: acuminate
Leaves: margin: denticulate, serrulate
Leaves: length (cm): 9-24 cm
Leaves: width (cm): 3.6-8 cm
Leaves: indumentum: glabrous, scaberulous
Venation: palmate-pinnate
Petioles: length (cm): 3-12 cm
Petioles: diameter (mm): 5 mm
Petioles: indumentum: glabrous
Bracts: caducous
Bracts: apex: mucronate, acuminate
Bracts: margin: entire
Bracts: length (mm): 19 mm
Bracts: width (mm): 12 mm
Bracts: indumentum: glabrous
Peduncles:: pendulous
Peduncles: length (cm): 2 cm
Peduncles: diameter (mm): 2 mm
Flowers: color: white
Male flower petals: number: four
Outer 2 tepals of male flowers: shape: broadly ovate
Outer 2 tepals of male flowers: length (mm): 8-13 mm
Outer 2 tepals of male flowers: width (mm): 6-12 mm
Inner 2 tepals of male flowers: shape: elliptic
Inner 2 tepals of male flowers: length (mm): 6-11 mm
Inner 2 tepals of male flowers: width (mm): 4-6 mm
Stamens: shape: clavate
Anthers: length (mm): 1.7-2.4 mm
Anthers: diameter (mm): 0.4-0.6 mm
Filaments: shorter than anthers
Filaments: length (mm): 0.8-1.4 mm
Female flower tepals: number: six
Female flower tepals: equality of size: equal, subequal
The largest tepals of female flowers: shape:: broadly obovate
Styles: number: three
Styles: length (mm): 4.9-5.4 mm
Ovary: shape: ellipsoid
Locules number: three
Fruits: typ: baccate
Fruits: wings: wingless
Seeds: shape: broadly ellipsoid
Seeds: length (mm): 0.25-0.28 mm
Seeds: diameter (mm): 0.18-0.21 mm
Seed apex: shape: rotund
Seed micropylar end: constricted
Chrosome numer 2n=: 22
Elevation (m): 50-2200 m
Distribution: Taipei, Taoyuan, Ilan, Hsinchu, Miaoli, Taichung, Nantou, Hualien
Plant: growth form: terrestrial
Phenology: evergreen
Stems: habit: with very short(or without) rhizomes from which upright stem arise
