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Murdannia taiwanensis Peng var. taiwanensis

台灣水竹葉

Perennial herbs. Roots fibrous, thickened, to 3.4 mm across. Stems dimorphic: the vegetative shoot acaulescent, erect; the flowering shoots one to several per plant, ascending. Rosette leaves linear, 11-24 cm long, 5-10 mm wide, apex acuminate, the midrib above the leaf white, glabrous; cauline leaves linear to lanceolate, 2.3-4.7 cm long, 4-5 mm wide; sheath glabrous, ciliate at mouth. Prophyll sheathing, 5-25 mm long, apex bifid. Inflorescence terminal, consisting of a single cincinnus, 1-3 flowering shoots from axils of the cauline leaves; bracts ovate to lanceolate, ca. 5 mm long, ca. 2.8 mm wide, fugacious, bracteoles ca. 3.8 mm long, ca. 2 mm wide; pedicels ca. 5 mm long. Flowers 14-18 mm in diam., dimorphic: in functionally male flowers the style central and straight, the pistil vestigial, ca. 2.4 mm long, stamens symmetrically arranged; in bisexual flowers the style curved to one side and stamens to the opposite side; sepals lanceolate, boat-shaped, greenish, equal, ca. 4.5 mm long, ca. 2.2 mm wide, apex obtuse; petals suborbicular to obovate, ca. 6-7.5 mm long, 5-5.5 mm wide, purple, veins conspicuous in fresh material. Fertile stamens 2, equal, antesepalous, anthers purple, narrowly elliptic-oblong, ca. 1.8 mm long; pollen whitish; filaments ca. 6 mm long, bearded, the hairs moniliform, purple; staminodes 4, unequal, one antesepalous, the anther vestigial, the filament ca. 4.4 mm long, bearded; the others antepetalous, equal, antherodes trilobed, yellow, ca. 1.2 mm long, filaments ca. 3.4 mm long; ovary ellipsoid in outline, acutely 3-angled, ca. 2 mm long; style thickened toward the base, ca. 5 mm long, glabrous. Capsule trilocular, 3-valved, ellipsoid, 3-angled, ca. 7 mm long, locules 2-seeded, apex mucronulate. Seed fulvous, deltoid in outline, 1.8-2 mm long, ca 1.4 mm across, apex acute, base truncate, dorsiventral, dorsal surface convex, ventral carinate, scrobiculate, tomentum, embryotega semilateral, hilum linear. Chromosome number: n = 28.

Pingtung: Kuraru, Matuda 456 (TAI); Chunjih Hsiang: Chi-Chia logging tract, elev. ca. 300-600 m, Wang & Yang 8921 (TNU, HAST); same loc., elev. ca. 760 m, Liu 725 (HAST). Kaoshiung: Liukuei to Meinung. On steap rocky slope face by a small rocky rivey bed, elev. ca. 200 m, Chen & Wang 331 (HAST); Liukuei to Meinung, Hsien Rd. #184, between Liukuei tunnel #2 and #3, Y. J. Chen et al. 794 (TNU); Liukuei Hsiang: West of Hwy 184, off Liukuei bridge No.5, in a rockey valley, elev. ca. 230 m, Peng 14766; Liukuei Hsiang: West of Hwy 184, milage sign 52 and 53 km, in a narrow valley, elev. ca. 240 m, Peng 14797 (HAST).

Endemic, in southern part of Taiwan, found in bluffs along valley. Flowering in May.- Nov.

本分類群形態上和M. angustifolia及M. loriformis極為類似,然而本種植物根粗,可達3.4 mm,葉子較厚,葉上表面中肋為白色,花極大,直徑可達1.8cm,花瓣上有紫色的脈,花瓣在盛開時常會向後展,退化雄蕊花絲上少有念珠狀毛,種子表面有密茸毛及小凹陷;M. angustifolia植株基部有球莖狀的莖,葉極狹,花瓣先端淺凹,退化雄蕊花絲上著生念珠狀的毛,至於種子形態則和本種植物十分類似;M. loriformis根細,寬小於2 mm,花徑小於1.4 mm,種子表面有密茸毛構成輻射紋或網紋,但沒有小凹陷。這三個分類群在製成標本後更加難以辨認,必須仔細比較才能區別。此三種之地理分布不同,M. loriformis廣泛分布於全省; M. angustifolia及本種植物分布於台灣南部,M. angustifolia則更侷限分布在恆春半島海岸邊。

Phenology: perennial herbs, evergreen

Roors form: roots fibrous, fleshy

Stems form: monomorphic stem, dimorphic stem

Monomorphic stems habit: acaulescent

Dimorphic stems habit: vegetative shoot acaulescent; flowering shoots ascending

Rooting at nodes or not: absent

Lateral shoots: not perforating

Phyllotaxis: spiral phyllotaxis

Leaves: petiole: sessile

Leaves: shape (if dimorphic leaves, selected by rosette leaves): linear

Leaves: apex: acuminate

Leaves: length (cm): 11-24

Leaves: width (cm): 0.5-1

Leaves indumentum: upper surface: glabrous

If upper surface glabrous, leaves indumentum: margin: glabrous

Prophyll: sheathing, apex bifid

Inflorescence: take plance: terminal

Inflorescence: rhipidium

Bracts: shape: lanceolate, ovate

Corolla: actinomorphic: petal 3, base free, equal or subequal

Flowers: bisexual, functionally male flowers

Sepals: sepal 3, equal

Sepals: color: greenish

Sepals: base: free

Sepals: shape: lanceolate

Sepals: indumentum: glabrous

Sepals: length (if dimorphic sepals, selected by anticous two) (mm): 4.5

Sepals: width (if dimorphic sepals, selected by anticous two) (mm): 2.2

Flowers: color: pink, purple, or blue

Petals: shape: suborbicular

Petals: length (mm): 6-7.5

Petals: width (mm): 5-5.5

Stamens form: trimorphic: fertile stamens 2

Ovarys: shape: ellipsoid

Ovarys: indumentum: glabrous

Ovarys: length (mm): 2

Styles: indumentum: glabrous

Styles: length (mm): 5

Capsules: dehiscence: dehiscent

Capsules: shape: ellipsoid

Capsules: trilocular

Capsules: indumentum: glabrous

Capsule locules seeds: number: 2

Capsules: length (mm): 7

Seeds: lengh (if dimorphic seeds, selected by dorsal one) (mm): 1.8-2

Seeds: width (if dimorphic seeds, selected by dorsal one) (mm): 1.4


 

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