Habit: tree

Sexuality: monoecious, dioecious

Branches: whorled

Leaves: Position: spirally arranged

Leaves: Shape: needle-like, broadly ovate, triangular

Microsporophylls: densely arranged

Seed Cones: Shape: ovoid, ellipsoid

Seed Cones: Bract scales: margin ridged, apex reflexed or upcurved or cuspidate, apex pointed

Seed scales: adnate with bracts in basal, detached in apical part

Seeds: Other Characteristics: with 2 lateral wings, connate with bracts

Cotyledons number: 2, 4

Germination: epigeal, hypogeal