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Athyriaceae | Family List | Athyriaceae

Athyrium Roth


Athyrium nigripes
Athyrium nigripes (Bl.) Moore

Credit: Liu, Yea-chen

Pseudocystopteris Ching

Terrestrial ferns; rhizome usually stout and erect, rarely long creeping; scales thin, lancrolate or ovate and light brown, some narrow and black . Stipe green, stramineous, or purple, with 2 vascular strands at base and uniting into U-shaped strand at middle rachies; lamina pinnate to decompound; veins all free; rachis and costae grooved and connected on dorsal side, some species with spines on the dorsal side of rachies, costae and costulate. Sori oblong or round, usually crossing a veinlet, some species with Diplazium type. Indusia linear, J-shape or reniform.

About 24 species in Taiwan, most species in Taiwan grown at altitude 1000 to 3000 m.

1 Rhizome long creeping   (2)
+ Rhizome erect   (5)
2 (1) Fronds bipinnatifid or more complex   (3)
+ Fronds pinnatifid or pinnate   (4)
3 (2) Fronds bipinnate or tripinnatifid   Athyrium nipponicum
+ Frond tripinnate or more complex   Athyrium atkinsonii
4 (2) With distinct apex pinnae, later pinnae with petiole   Athyrium cumingianum
+ Without distinct apex pinnae, later pinnae without petiole   Athyrium sheareri
5 (1) Without spines on rachis, costae and costulate   (6)
+ Spines on rachis, costae or costulate   (13)
6 (5) Pinnate, or with free pinnulate in the base of pinnae   (7)
+ Bipinnate or more complex   (9)
7 (6) Fronds glabrous   Athyrium anisopterum
+ Frond pubescent, densely covered with unicellular hairs   (8)
8 (7) Margin of pinnae entire or crenate, with obviouly auricaulate   Athyrium nakanoi
+ Margin of pinnae serrate, sometimes with free pinnulate in the base of pinnae   Athyrium australlium
9 (6) Bipinnatifid, fronds covered gland hairs, most pinnae reflex   Athyrium reflexipinnum
+ Bipinnate or more complex   (10)
10 (9) Sori short straight, less variable   (11)
+ Sori reniform, or variable   (12)

  • List of lower taxa


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