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Athyriaceae | Family List | Athyriaceae | Athyrium

Athyrium arisanense (Hay.) Tagawa, Acta Phytotax. Geobot. 2(3). 195. 1933.


Athyrium arisanense
Athyrium arisanense var. kuratae

Credit: Liu, Yea-chen

Diplazium arisanense Hayata in Icon. Pl. Formosan. 4: 212-213, f. 144. 1914.

Terrestrial; rhizome errect, sometimes ascending; Stipe length 5-40 cm, green, stramineous or purple; scales on stipe base dark brown or black, concolor, entire in edge; fronds bipinnatifid, pinnae 7-25 by 3-10 cm, end of pinnae caudate or acute; rachis and costae pubescent at abaxil surface, cover densely short unicellular hairs and some long-soft hairs and gland hairs; lowest pinnae sessile or with less 1 mm petiole; pinnulate opposite or near opposite, length 0.5-5 cm, acroscopic and basiscopic pinnae equal length, excepted some lowest pinnae with basiscopic enlongate pinnulate; serrate in edge; sori short linear or hook crossing over the veinlet, sometimes two sides in veinlet (Diplazium type), indusia shape as sori, edge entire or crisped; sori position at middle veinlet, not attach to the costae and costulate, 1 to 5 pairs in one pinnulate, length 1-3 mm; spore yellow brown, ellipsoidal, surface cover flat dropped perispore.

地生; 根莖直立, 偶倒臥; 葉柄長5-40公分, 綠色, 草桿色或紫紅色; 葉柄基部鱗片深褐色或黑色, 邊緣全緣, 具或不具黃色窄邊; 二回羽狀深裂; 一回羽葉長7-25公分, 寬3-10公分, 末端具漸尖至長尾尖; 葉軸及羽軸遠軸面均具短, 單細胞腺毛, 以及多細胞長軟毛和末端膨大腺毛; 最下方一至三對一回羽葉基部小羽片具極短或不具柄, 近對生; 其他部位小羽片基部收縮, 不具柄; 二回小羽片末端圓鈍或尖頭, 長0.5-5公分, 羽軸兩側羽片長度相近, 僅最下方羽片的下方小羽片延長成鐮刀狀,或與上方小羽片相同, 邊緣具斜展的鋸齒; 孢子囊群短直, 以及少數彎勾狀或背生於小脈兩側, 孢膜與孢子囊群同型, 邊緣全緣或具波浪緣; 孢子囊群位於小脈中段, 不緊靠羽軸及小羽軸, 每一小羽片一至多對, 長度1-3公釐; 孢子黃褐色, 長橢圓形, 表面具可剝落的扁平皺紋.

Holotype: Taiwan: Jan. 1912, B. Hayata & S. Sasaki s.n. (HT: ?)


South China, Taiwan, and Japan.

This species is much similar with Athyrium kuratae Serizawa and Athyrium kenzosatakei Kurata. It also similar with Athyrium hainanense Ching but the later one without spine on adaxil costae.

Kurata, S. 1961. On the Japanese ferns belonging to the Athyrium otophorum group. Science report of the Yokosuka city museum 6:7-26, pl. 2-3.
Zhang, X. C. 1996. On the genus Athyrium ser. epiraches Ching et Y. T. Hsieh ex Y. T. Hsieh. Acta Phytotaxonomica Sinica 34(2): 180-193.

Habitat: Edge of forest or side by the trail, Understory of Forest

Leaf shape: Broad-Lanceolate, Deltoid

Apex of Leaf: Acuminate

Leaf Division: Bipinnatifid, Bipinnate

Length of Stipe: 20 (10-60)

Length of Lamina: 30 (10-60)

Width of Lamina: 30 (15-40)

Stipe Color: Purple, Purple to Stramineous, Stramineous, Green

Free Pinnulate in basal Pinnae: Yes, No

Form of basal Pinnulate arrangement: Opposite

Apex of Pinnae: Acuminate

Petioled in pinnae: Short Petiole (less than 3 mm), Sessile

Length of Lowest Pinnae: 15 (10-20)

Wideth of Lowest Pinnae: 7 (5-10)

Lowest pinnae with enlarged basiscopic pinnules (more than 3 mm): Yes, more than 3 mm

Apex of terminal fragment: Round or Obtuse, Acuminate

Margin of terminal fragment: Crenate, Serrulate

Type of Basal of Pinnule: Broadly adnate, Constricted

Anterior pinna or pinnule auriculate: not auricaulate

Lowest Pinnae with shorter basal pinnule: Yes, basal pinnule shorter than next two

Second Lowest Pinnae with shorter basal pinnule: Yes, Basal Pinnulate shorter than next two

Abaxial Surface of Rachis with Unicellular Hairs ?: Yes, densely coved with unicellular hairs (pubescent)

Abaxial Surface of Rachis with Gland Hairs?: Yes, there are rare gland hairs on rachis

Abaxial Surface of Coaste with Unicellular Hairs ?: Yes, densely coverd unicellular hairs on costae (pubescent)

Type of Adaxial Costae Spines: Short spines

Spines Existing on Adaxial Costulate: No, spines Absent on costulate

Type of Scales on Basal Stipe: Lanceolate

Color of Scales on Basal Stipe: Dark Brown to Black

With Adventitious Bud near the Apex of Leave: No, adventitious buds absent

Sori Shape: Curved, Diplazium type

Sori Position: Middle of veinlet

Spore Surface Type: Rugate Type

Type of Rhizome: Short Erect

Geographic Distribution: Taiwan - Japan

Altitude: 1800 (900-2500)


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  • Athyrium arisanense (Hayata) Tagawa (Liu, Yea-chen)
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  • Athyrium arisanense var. kuratae (Liu, Yea-chen)
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  • Original Publish Data
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    Flora of China  
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