Distribution: Endemic to Madagascar

Bioclimate: Humid

Vegetation Formation: Forest, Mangrove, Freshwater Wetland, Saline Wetland

Elevation (m): 0-499, 500-999

Province: Antsiranana, Fianarantsoa, Toamasina

Protected Areas : Mananara-Nord, Masoala, Ranomafana

Preliminary Conservation Assessment : A1cd+2cd

Note: Some specimens collected from higher elevation need to be checked ( Ranomafana, Ambatovy, Sahananto - 950m - 1200m)

Updated on: 11 Sep. 2006