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8. Actinodaphne Nees in Wallich, Pl. Asiat. Rar. 2: 61, 68. 1831.

黄肉楠属 huang rou nan shu

Authors: Puhua Huang & Henk van der Werff

Evergreen trees or shrubs, dioecious. Leaves usually clustered or nearly verticillate, rarely alternate or opposite, unlobed, pinninerved, rarely triplinerved. Umbels solitary or clustered or arranged in a panicle or raceme; involucral bracts imbricate, caducous. Perianth tube short; perianth segments usually 6 in 2 whorls of 3 each, nearly equal, rarely persistent. Flowers unisexual. Male flowers: fertile stamens usually 9 in 3 whorls of 3 each; filaments of 1st and 2nd whorls eglandular, of 3rd whorls 2-glandular at base; anthers all introrse and 4-celled, cells opening by lids; rudimentary pistil small or lacking. Female flowers: staminodes as many as stamens of male flowers; ovary superior; stigma shield-shaped or dilated. Fruit seated on shallow or deep cup-shaped or discoid perianth tube.

About 100 species: tropical and subtropical regions of Asia; 17 species (13 endemic) in China.

Actinodaphne sessilifructa C. J. Qi & K. W. Liu (Guihaia 7: 215. 1987) was described from SW Hunan (Tongdao) but could not be treated here because no material was seen by the present authors.


1 Leaf blade triplinerved   (2)
+ Leaf blade pinninerved   (3)
       
2 (1) Buds, branchlets, and leaf blade densely ferruginous pubescent; fruit 2.5-4.5 × 1-2 cm, fruiting pedicel robust, 0.5-0.6 cm × 3-4 mm.   1 A. obovata
+ Buds, branchlets, and leaf blade glabrous; fruit ca. 2.5 × 1.5 cm, fruiting pedicel slender, ca. 1 cm × 2 mm.   2 A. menghaiensis
       
3 (1) Bud scales persistent, surrounding bases of branchlets   (4)
+ Bud scales caducous   (9)
       
4 (3) Bud scales smaller, 3-8 × 2-6 mm, densely arranged   (5)
+ Bud scales larger, 10-22 × 6-10 mm, sparsely arranged   (6)
       
5 (4) Leaf blade narrowly lanceolate, thickly leathery, lateral veins 18-26 pairs, obscure or nearly obscure abaxially; fruit glabrous.   3 A. obscurinervia
+ Leaf blade oblanceolate or oblong, leathery, lateral veins 6-10 pairs, conspicuously prominent abaxially; fruit yellow appressed tomentose.   4 A. trichocarpa
       
6 (4) Petiole less than 7 mm; lateral veins 7-9 pairs; leaf base rounded or broadly cuneate.   5 A. koshepangii
+ Petiole more than 11 mm; lateral veins 11-15 pairs; leaf base cuneate   (7)
       
7 (6) Petiole glabrous; leaf blade glabrous abaxially (only gray puberulent when young, soon glabrous).   6 A. omeiensis
+ Petiole and leaf blade abaxially densely tomentose   (8)
       
8 (7) Leaf blade elliptic, oblong-elliptic, or obovate-elliptic, broader, mostly more than 5 cm in diam.; petiole up to 4 cm; fruit subglobose, fruiting pedicel 4-5 mm.   7 A. kweichowensis
+ Leaf blade elliptic-lanceolate, narrower, mostly less than 5 cm in diam.; petiole up to 2 cm; fruit oblong, fruiting pedicel 11-15 mm.   8 A. forrestii
       
9 (3) Lateral veins 30-40 pairs or more.   9 A. lecomtei
+ Lateral veins less than 15 pairs   (10)
       
10 (9) Inflorescence or fruiting inflorescence paniculiform   (11)
+ Inflorescence or fruiting inflorescence umbellate   (12)
       
11 (10) Branchlets, young leaves, and inflorescence ferruginous tomentose; leaf blade obovate, rarely elliptic; fruiting tube subflat disciform.   15 A. pilosa
+ Branchlets with dense grayish appressed tomentum, leaf blade pubescent along midrib and veins abaxially, and inflorescence white sericeous; leaf blade lanceolate; fruiting tube shallowly cup-shaped.   14 A. henryi
       
12 (10) Young branchlets glabrous.   16 A. acuminata
+ Young branchlets hairy   (13)
       
13 (12) Petiole shorter, 3-8 mm   (14)
+ Petiole longer, usually 10-40 mm   (16)
       
14 (13) Leaf blade oblanceolate or obovate-lanceolate; filaments longer, ca. 6.5 mm, villous.   10 A. tsaii
+ Leaf blade not as above; filaments shorter, ca. 4 mm, glabrous   (15)
       
15 (14) Leaf blade oblong-ovate, base rounded or broadly cuneate, lateral veins 7-9 pairs.   5 A. koshepangii
+ Leaf blade oblong-lanceolate to oblong, base acuminate or acute, lateral veins 8-13 pairs.   11 A. cupularis
       
16 (13) Leaf blade oblanceolate, obovate-lanceolate, occasionally oblong, thinner   (17)
+ Leaf blade long lanceolate, lanceolate, elliptic to oblong-elliptic, thicker   (18)
       
17 (16) Leaves 5-7-verticillate, leaf blade 14-20.5 × 3.5-6.5 cm, appressed pubescent abaxially; fertile stamens 9-15, filaments glabrous.   12 A. paotingensis
+ Leaves scattered, leaf blade 6-12 × 1.8-3 cm, densely gray-yellow villous abaxially; fertile stamens 9, filaments villous.   17 A. mushaensis
       
18 (16) Midrib prominent on both surfaces of leaf blade; fruit globose, smaller.   13 A. glaucina
+ Midrib concave on leaf blade adaxially; fruit oblong or subglobose   (19)
       
19 (18) Leaf blade abaxially and petiole glabrous; fruit subglobose, to 20 mm in diam..   6 A. omeiensis
+ Leaf blade abaxially and petiole hairy; fruit oblong, 6-8 mm in diam..   8 A. forrestii

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