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FOC | Family List | FOC Vol. 12 | Theaceae

9. Adinandra Jack, Malayan Misc. 2(7): 49. 1822.

杨桐属 yang tong shu

Shrubs or trees, evergreen. Leaves petiolate; leaf blade margin entire or serrulate. Flowers bisexual, solitary or paired. Pedicel usually recurved, seldom erect; bracteoles 2, inserted at apex of pedicel, alternate or opposite, persistent or caducous. Sepals 5, persistent, imbricate, unequal, thick, concave, basally slightly connate. Petals 5, imbricate, outside glabrous or sericeous, basally connate. Stamens 15-60, in 1-5 whorls; filaments connate or distinct, adnate to base of sepals, glabrous or with pubescence; anthers basifixed, oblong, with filiform trichomes, connective projected into an apiculum. Ovary pubescent or glabrous, (2 or)3-5-loculed, placentation axile; ovules 20-100 per locule; style l, simple or apically 3-5-lobed. Fruit baccate, many-seeded, with a persistent style. Seeds brown, minute, foveolate to pitted, shiny; embryo inflexed; endosperm abundant; cotyledons semiterete.

About 85 species: Bangladesh, Cambodia, China, India, Indonesia, S Japan, Laos, Malaysia, Myanmar, New Guinea, Philippines, Sri Lanka, Thailand, Vietnam; tropical Africa; 22 species (17 endemic) in China.

Adinandra stenosepala Hu is the basionym of Xantolis stenosepala (Hu) P. Royen in the Sapotaceae (see Fl. China 15: 209. 1996).


1 Style apically parted or lobed   (2)
+ Style simple   (6)
       
2 (1) Ovary pubescent.   6 A. filipes
+ Ovary glabrous   (3)
       
3 (2) Style villous.   16 A. elegans
+ Style glabrous   (4)
       
4 (3) Style apically slightly 2-lobed.   19 A. retusa
+ Style apically 3- or 4-lobed   (5)
       
5 (4) Leaf blade abaxially reddish glandular punctate; pedicel 0.6-0.8 cm.   17 A. pingbianensis
+ Leaf blade abaxially not glandular punctate; pedicel 1-2 cm.   18 A. nitida
       
6 (1) Ovary glabrous   (7)
+ Ovary pubescent   (11)
       
7 (6) Leaf blade 4-7 mm wide.   A. angustifolia
+ Leaf blade 10 mm wide or more   (8)
       
8 (7) Ovary more than 3-loculed   (9)
+ Ovary 3-loculed   (10)
       
9 (8) Petiole 10-20 mm, leaf blade elliptic, 13-18 × 5.5-8.5 cm, abaxially glabrous, base broadly cuneate, apex acute and with an obtuse tip; pedicel 1-1.8 cm.   4 A. grandis
+ Petiole 1-1.5 mm, leaf blade lanceolate to narrowly lanceolate, 7.5-12.5 × 1.9-2.5 cm, abaxially appressed pubescent and densely so along midvein, base obliquely auriculate, apex long acuminate to caudate-acuminate; pedicel 0.5-0.7 cm.   5 A. auriformis
       
10 (8) Petiole 0.3-0.5 cm; leaf blade abaxially not glandular; petals ovate to suborbicular, outside sericeous; filaments glabrous.   20 A. howii
+ Petiole 1.5-2 cm; leaf blade abaxially glandular punctate; petals broadly obovate to oblong-obovate, outside glabrous; filaments with dense filiform trichomes on apical 1/2.   21 A. nigroglandulosa
       
11 (6) Ovary 5(or 6)-loculed   (12)
+ Ovary 3-loculed   (15)
       
12 (11) Style glabrous.   3 A. latifolia
+ Style sericeous   (13)
       
13 (12) Leaf blade abaxially reddish brown glandular punctate.   2 A. hainanensis
+ Leaf blade abaxially not glandular punctate   (14)
       
14 (13) Leaf blade secondary veins 20-26 on each side of midvein; bracteoles oblong, 6-7 mm; sepal apex obtuse; petals 10-13 × 6-7 mm.   1 A. megaphylla
+ Leaf blade secondary veins 12-16 on each side of midvein; bracteoles ovate to long ovate, 2-3.5 mm; sepal apex acute; petals 8-10 × 5-6 mm.   7 A. integerrima
       
15 (11) Style pubescent at least basally (A. hirta var. macrobracteata sometimes with a glabrous style)   (16)
+ Style glabrous   (20)
       
16 (15) Leaf blade abaxially hirsute with trichomes projecting beyond margin   (17)
+ Leaf blade abaxially appressed pubescent with trichomes not projecting beyond margin   (18)
       
17 (16) Petiole 8-10 mm; pedicel 6-16 mm, bracteoles caducous; petals outside hirtellous along median portion; style densely hirsute on basal 2/3 or entire length.   10 A. glischroloma
+ Petiole 5-7 mm; pedicel 5-6 mm, bracteoles persistent; petals outside glabrous; style hirtellous or only basally hirtellous.   11 A. hirta
       
18 (16) Pedicel 2-4 cm.   6 A. filipes
+ Pedicel 0.5-0.7 cm   (19)
       
19 (18) Petiole 3-5 mm; leaf blade acuminate; bracteoles caducous; filaments 1.5-2 mm, glabrous; style ca. 10 mm.   8 A. lasiostyla
+ Petiole 8-15 mm; leaf blade abruptly acute; bracteoles persistent; filaments 2-4 mm, pubescent; style 3-4 mm.   9 A. epunctata
       
20 (15) Sepals 4-6 mm; petals outside densely sericeous along median portion   (21)
+ Sepals 7-11 mm; petals outside glabrous   (22)
       
21 (20) Petiole 2-3 mm; leaf blade lanceolate to oblong-lanceolate, base broadly cuneate to rounded; pedicel 0.5-0.9 cm; petals lanceolate to obovate-lanceolate, apex acuminate; stamens 15-17; persistent sepals reflexed.   14 A. lancipetala
+ Petiole 5-7 mm; leaf blade oblong to oblong-ovate, base cuneate; pedicel 1-2 cm; petals ovate, apex rounded and mucronate; stamens 25-30; persistent sepals not reflexed.   15 A. bockiana
       
22 (20) Leaf blade margin apically serrate; pedicel 2.5-3.5 cm.   12 A. formosana
+ Leaf blade margin entire; pedicel 0.5-2 cm   (23)
       
23 (22) Petiole 5-7 mm; leaf blade 9-12.5(-16) × 3-4.5(-5.8) cm; pedicel 5-6 mm, bracteoles 6-10 mm.   11 A. hirta
+ Petiole 3-5 mm; leaf blade 4.5-9 × 2-3 cm; pedicel ca. 20 mm, bracteoles 2-3 mm.   13 A. millettii

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