|
6. Ajuga Linnaeus, Sp. Pl. 2: 561. 1753.
筋骨草属 jin gu cao shu
Plants annual, biennial or perennial, herbaceous, rarely shrubs. Leaves simple; leaf blade papery, margin dentate to incised, rarely subentire. Verticillasters 2- to many flowered, in false spikes; floral leaves similar to stem leaves or gradually reduced to bracts, rarely dissimilar, larger than stem leaves. Flowers subsessile. Calyx ovoid to globose, campanulate to funnelform, 10-veined, sometimes with inconspicuous accessory veins; teeth 5, slightly irregular. Corolla purple to blue, rarely yellow or white, 2-lipped, often persistent in fruit; tube straight to slightly curved, base slightly bent/swollen; throat slightly dilated, villous annulate, rarely glabrous inside; upper lip straight, entire to 2-lobed; lower lip elongate, 3-lobed, with middle lobe obcordate to nearly flabellate and lateral lobes oblong. Stamens 4, didynamous, exserted from upper lip, involute in bud, anterior 2 longer; filaments straight to slightly curved; anther cells 2, apically confluent. Style subequally 2-cleft, lobes subulate. Nutlets obovoid, triquetrous, netted on back, lateral-ventral side with an areole 1/2-2/3 its length, with an elaiosome.
About 40-50 species: Asia, Europe, especially in the Near East; 18 species in China.
|
|
|
|
| 1 |
Verticillasters 2(-4, rarely more)-flowered, widely spaced; floral leaves similar to stem leaves; fertile branches (only) erect to ascending. |
|
(2) |
| + |
Verticillasters 6- or more flowered, close together; floral leaves unlike stem leaves (except in A. linearifolia); plants erect, rarely trailing |
|
(5) |
| |
|
|
|
| 2 (1) |
Base of corolla tube not villous annulate inside; leaf base cordate to subtruncate |
|
1 Ajuga lobata |
| + |
Base of corolla tube villous annulate inside; leaf base cuneate to subtruncate, decurrent. |
|
(3) |
| |
|
|
|
| 3 (2) |
Leaves spatulate, 2-3 × 4-6 mm |
|
2 Ajuga pygmaea |
| + |
Leaves ovate to oblong, more than 10 mm wide. |
|
(4) |
| |
|
|
|
| 4 (3) |
Leaves 3-5.3 × 2.5-3.8 cm, ovate to ovate-oblong, basally cuneate to subtruncate, slightly decurrent |
|
3 Ajuga sciaphila |
| + |
Leaves 2-3.4 × 1.2-2.3 cm, oblong to ovate-oblong, basally cuneate to decurrent |
|
4 Ajuga nubigena |
| |
|
|
|
| 5 (1) |
Floral leaves longer than flowers, whitish yellow, white, or purple |
|
5 Ajuga lupulina |
| + |
Floral leaves equal to or slightly shorter than flowers, rarely longer, green or purplish |
|
(6) |
| |
|
|
|
| 6 (5) |
Corolla tube (1.5–)2–3 cm, upper lip 2-lobed |
|
6 Ajuga ovalifolia |
| + |
Corolla tube less than 1.6 cm, rarely to 2 cm, upper lip shallowly 2-lobed |
|
(7) |
| |
|
|
|
| 7 (6) |
Floral and stem leaves similar, of 1 color; stem leaves linear to linear-lanceolate |
|
18 Ajuga linearifolia |
| + |
Floral and stem leaves dissimilar; stem leaves of other shapes |
|
(8) |
| |
|
|
|
| 8 (7) |
Corolla tube erect to slightly curved, not saccate or bent above annulus of hairs |
|
(9) |
| + |
Corolla tube slightly swollen, shallowly saccate to bent above annulus of hairs |
|
(14) |
| |
|
|
|
| 9 (8) |
Calyx ciliate along veins and teeth margin, villous to puberulent apically, glabrous basally, or subglabrous throughout |
|
(10) |
| + |
Calyx strigose, villous, or lanate-villous throughout, especially on teeth |
|
(12) |
| |
|
|
|
| 10 (9) |
Stems 25–40 cm tall, purplish red or greenish purple, glabrous, gray villous only on younger parts; leaves ovate-oblong to narrowly oblong, 4–7.5?×?3.2–4 cm, irregularly double dentate |
|
7 Ajuga ciliata |
| + |
Stems less than 20 cm tall, sometimes to 30 cm, not brightly colored, densely gray pubescent to gray villous; leaves ovate to lanceolate-oblong, less than 3.2 cm wide, margin repand |
|
(11) |
| |
|
|
|
| 11 (10) |
Inflorescences 1–3 cm; floral leaves usually shorter than flowers; calyx white villous on teeth |
|
9 Ajuga campylanthoides |
| + |
Inflorescences ca. 6 cm; floral leaves much longer than flowers; calyx ciliate along veins and apical teeth margin |
|
11 Ajuga forrestii |
| |
|
|
|
| 12 (9) |
Calyx strigose; corolla tube slightly curved; stamens nearly included; plants subprostrate, basally stoloniferous |
|
10 Ajuga campylantha |
| + |
Calyx villous or lanate-villous; corolla tube straight; stamens slightly exserted; plants erect or stoloniferous |
|
(13) |
| |
|
|
|
| 13 (12) |
Stems erect, unbranched, 6–21 cm tall, densely gray lanate or silky |
|
8 Ajuga multiflora |
| + |
Plants with stolons and erect fertile stems ca. 10 cm tall, gray villous or lanate-villous |
|
12 Ajuga bracteosa |
| |
|
|
|
| 14 (8) |
Calyx teeth lanceolate to lanceolate-triangular; plants spreading ascending or with stolons |
|
13 Ajuga pantantha |
| + |
Calyx teeth narrowly triangular to broadly ovate; plants erect, sometimes trailing-ascending |
|
(15) |
| |
|
|
|
| 15 (14) |
Corolla 6–8(–9) mm; calyx teeth ovate to ovate-triangular; stamens exserted ca. 1/2 their length |
|
(16) |
| + |
Corolla more than 8 mm; calyx teeth narrowly or short triangular; stamens exserted 1/4–1/3 their length |
|
(17) |
| |
|
|
|
| 16 (15) |
Corolla white, upper lip shorter than lateral lobes of lower lip; calyx teeth ovate-triangular |
|
16 Ajuga dictyocarpa |
| + |
Corolla blue, blue-purple, or purple, upper lip nearly as long as lateral lobes of lower lip; calyx teeth ovate to broadly ovate |
|
17 Ajuga macrosperma |
| |
|
|
|
| 17 (15) |
Leaves spatulate to suboblong; plants prostrate, with basal leaves at anthesis |
|
14 Ajuga decumbens |
| + |
Leaves broadly oblong to obovate-elliptic; plants erect or rarely prostrate, branched at base, without basal leaves at anthesis |
|
15 Ajuga nipponensis |
|
|
|
List of lower taxa
| Related Links (opens in a new window) |
|
Treatments in Other Floras @ www.efloras.org
Other Databases
|
|
|
|
|
|
|