3. Anisocampium C. Presl, Epimel. Bot. 58. 1851.
安蕨属 an jue shu
Authors: Zhongren Wang & Masahiro Kato
Kuniwatsukia Pichi Sermolli; Microchlaena Ching (1938), not Wight & Arnott (1833).
Plants terrestrial, small to medium-sized. Rhizomes long creeping, erect, or shortly creeping with ascending apex, clothed with brown lanceolate or linear scales. Fronds distant or caespitose; stipe usually stramineous, long, with sparse scales at base, upper part glabrate, grooved adaxially, groove open to rachis groove; lamina pinnate, oblong-lanceolate, ovate-oblong, or deltoid-ovate; pinnae pinnatifid at apex or terminal pinna similar to lateral pinnae; basal pinnae opposite or subopposite, shortly stalked; upper pinnae alternate, falcate or ascending, lanceolate, margin shallowly lobed, segments toothed, acuminate. Veins pinnate, lateral veins 3-5 pairs, simple or occasionally forked, free or connected to form 1 or 2 pairs of rhomboid areoles. Lamina papery when dried, glabrate adaxially, costae and midribs abaxially with brown linear-lanceolate small scales and short pale hairs. Sori orbicular, medial or subbasal on veins; indusia small, orbicular-reniform, membranous, ciliate at margin, fugacious. Spores bilateral, perispore surface with lophate ridges. x = 40.
Four species: tropical and subtropical regions of SE Asia and temperate areas of E Asia; four species in China.
A recent molecular and systematic study shows that Anisocampium includes Kuniwatsukia and Athyrium niponicum and is basal in the Athyrium complex (Liu et al. Taxon 60: 824-828. 2011).