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FOC | Family List | FOC Vol. 2-3 | Athyriaceae

3. Anisocampium C. Presl, Epimel. Bot. 58. 1851.

安蕨属 an jue shu

Authors: Zhongren Wang & Masahiro Kato

Kuniwatsukia Pichi Sermolli; Microchlaena Ching (1938), not Wight & Arnott (1833).

Plants terrestrial, small to medium-sized. Rhizomes long creeping, erect, or shortly creeping with ascending apex, clothed with brown lanceolate or linear scales. Fronds distant or caespitose; stipe usually stramineous, long, with sparse scales at base, upper part glabrate, grooved adaxially, groove open to rachis groove; lamina pinnate, oblong-lanceolate, ovate-oblong, or deltoid-ovate; pinnae pinnatifid at apex or terminal pinna similar to lateral pinnae; basal pinnae opposite or subopposite, shortly stalked; upper pinnae alternate, falcate or ascending, lanceolate, margin shallowly lobed, segments toothed, acuminate. Veins pinnate, lateral veins 3-5 pairs, simple or occasionally forked, free or connected to form 1 or 2 pairs of rhomboid areoles. Lamina papery when dried, glabrate adaxially, costae and midribs abaxially with brown linear-lanceolate small scales and short pale hairs. Sori orbicular, medial or subbasal on veins; indusia small, orbicular-reniform, membranous, ciliate at margin, fugacious. Spores bilateral, perispore surface with lophate ridges. x = 40.

Four species: tropical and subtropical regions of SE Asia and temperate areas of E Asia; four species in China.

A recent molecular and systematic study shows that Anisocampium includes Kuniwatsukia and Athyrium niponicum and is basal in the Athyrium complex (Liu et al. Taxon 60: 824-828. 2011).

1 Lamina pinnate, apex of lamina pinnatifid, not similar to lateral pinnae   (2)
+ Lamina terminated by a pinna similar or subsimilar to lateral pinnae   (3)
2 (1) Lamina pinnate throughout; pinnae usually shallowly lobed; sori orbicular.   1 A. sheareri
+ Lamina bipinnate; pinnules free, lobed; sori oblong, hooked, J-shaped, or horseshoe-shaped.   2 A. niponicum
3 (1) Pinnae up to 7 pairs, lanceolate, base cuneate or rounded; veins anastomosing.   3 A. cumingianum
+ Pinnae 10 pairs or more, linear, base truncate; veins free.   4 A. cuspidatum

Lower Taxa


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