1 |
Rhizome slender and long creeping; lamina herbaceous, rather thin and usually soft; axes and main veins adaxially with pale gray unicellular acicular hairs |
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(2) |
+ |
Rhizome usually stout, creeping to ascending, or erect; lamina papery to leathery, or rarely herbaceous especially when young; axes and main veins adaxially glabrescent or with scales, but not unicellular hairs |
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(4) |
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2 (1) |
Indusium entire; main veins abaxially with broadly ovate and bullate-based scales; stipe and primary rachis with copious scales. |
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1 A. miqueliana |
+ |
Indusium ciliate or entire; main veins abaxially with narrowly lanceolate, or hairlike and flat-based scales; upper part of stipe and primary rachis glabrescent |
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(3) |
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3 (2) |
Lamina pentagonal, 4(or 5)-pinnate, upper parts conspicuously abruptly narrowed toward apex and acuminate, brownish green when dried. |
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2 A. quadripinnata |
+ |
Lamina deltoid-pentagonal, 3-pinnate-pinnatifid, upper parts long acuminate, yellowish green when dried. |
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3 A. sinomiqueliana |
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4 (1) |
Lamina 4- or 5-pinnate, ultimate segments very narrowly lanceolate, with a single vein; sori solitary in each segment; indusia very firm, dark and entire, covering sorus when young and splitting into irregular segments when mature. |
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4 A. superba |
+ |
Lamina usually 2-4(or 5)-pinnate, rarely simply pinnate, ultimate segments asymmetrically oblong or rhomboid, with more than one vein; sori often more than one in each segment; indusia membranous, papery or firm, persistent or fugacious, margin entire or erose |
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(5) |
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5 (4) |
Lamina oblong-lanceolate, simply pinnate; pinnae lanceolate; free ultimate segments (pinnae) nearly symmetrically cuneate at base |
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(6) |
+ |
Lamina deltoid, ovate, or pentagonal, 2-4(or 5)-pinnate; pinnae usually deltoid or ovate-lanceolate; free ultimate segments usually asymmetrical at base with acroscopic auricles |
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(8) |
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6 (5) |
Stipe with dark brown scales at base; pinnae rachis abaxially with sparse hairlike scales; sori mostly dorsal at or below middle of veinlets. |
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5 A. neopodophylla |
+ |
Stipe with reddish brown scales at base; pinnae rachis abaxially glabrescent; sori terminal or subterminal on veinlets |
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(7) |
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7 (6) |
Lamina acuminate at apex or with a distinct terminal pinna; pinnae with aristate-tipped cartilaginous coarse teeth on margin. |
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6 A. blinii |
+ |
Lamina with a distinct terminal pinna; pinnae with non-aristate-tipped crenulate teeth on margin. |
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7 A. tsiangiana |
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8 (5) |
Sori dorsal on veinlets; ultimate segments with margin dentate or acutely serrate, never with aristate tips; texture thickly leathery. |
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8 A. cavaleriei |
+ |
Sori terminal on veinlets (or subdorsal but mixed with terminal ones on same lamina); ultimate segments with margin acutely dentate or pinnatilobate, teeth with or without aristate tips; texture papery, subleathery, or herbaceous |
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(9) |
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9 (8) |
Rhizome and base of stipe with shiny castaneous, ovate-lanceolate, and stiffly thickly papery scales; stipe upward and rachis throughout glabrescent and glossy |
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(10) |
+ |
Rhizome and base of stipe with blackish, grayish, or reddish brown, ovate-lanceolate, lanceolate, linear-lanceolate, or filiform, and membranous or thinly papery scales; stipe upward and rachis throughout ?scaly and often not glossy |
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(11) |
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10 (9) |
Lamina caudate toward apex, and with a distinct terminal pinna; lateral pinnae also caudate at apices. |
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9 A. amoena |
+ |
Lamina abruptly narrowed toward apex and caudate-acuminate, without distinct terminal pinna; lateral pinnae also caudate-acuminate at apices. |
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10 A. tonkinensis |
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11 (9) |
Rhizome ascending, with tufted, reddish brown, long, tortuous, filiform scales extending to stipe base |
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(12) |
+ |
Rhizome long to shortly creeping or ascending, often bearing grayish or blackish brown, linear-lanceolate, ovate-lanceolate, or subulate scales extending to stipe base |
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(16) |
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12 (11) |
Lamina 2-pinnate-pinnatifid, free ultimate segments 5-8 ?ca. 1.5 cm |
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(13) |
+ |
Lamina 3-5-pinnate, free ultimate segments at most to 2 ?1 cm |
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(14) |
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13 (12) |
Stipe base with only long hairlike scales; lamina leathery, lustrous and brownish when dried. |
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11 A. grossa |
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Stipe base with broad, denticulate-margined scales and with hairlike scales adaxially; lamina papery, dull dark green when dried. |
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12 A. fengii |
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14 (12) |
Fronds large, usually reaching ca. 1.7 m; lamina 5-pinnate; stipe with both subulate scales and broadly lanceolate, denticulate-margined thicker scales along both sides of adaxial groove at lower part above base. |
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15 A. gigantea |
+ |
Fronds medium-sized, at most to 1.5 m; lamina 3- or 4-pinnate; stipe with scales similar to those on rhizome or bearing slightly wider scales along both sides of adaxial groove near base |
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(15) |
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15 (14) |
Lamina papery, 3- or 4-pinnate; ultimate segments obviously separate from each other, apex and lobes mucronate, dull brownish when dried; indusia usually denticulate on margin. |
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13 A. globisora |
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Lamina thinly leathery, 4-pinnate; ultimate segments contiguous to imbricate, apex and lobes with non-aristate sharp teeth, shiny green when dried; indusia often entire on margin. |
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14 A. spectabilis |
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16 (11) |
Rhizome lignified, together with base of stipe bearing linear-lanceolate, lanceolate, or subulate, and thicker and firmer scales |
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(17) |
+ |
Rhizome succulent, especially at growing apex, bearing broadly lanceolate or ovate-lanceolate and more delicate scales extending to stipe base |
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(28) |
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17 (16) |
Fronds dimorphic; fertile lamina strongly contracted with much shorter and narrower pinnae and pinnules. |
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16 A. hainanensis |
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Fronds monomorphic, or with minor differences between fertile and sterile laminae (or pinnae on same lamina) |
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(18) |
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18 (17) |
Lamina abruptly acuminate to caudate-acuminate toward apex, most often with a conform terminal pinna resembling lateral pinnae |
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(19) |
+ |
Lamina gradually acuminate to strongly acuminate, gradually reduced without a conform terminal pinna |
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(21) |
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19 (18) |
Lamina 2- or 3-pinnate, with an elongated caudate-acuminate apex, lateral pinnae often linear-lanceolate (only lowest pinnae deltoid-lanceolate). |
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19 A. longipinna |
+ |
Lamina (2 or)3- or 4-pinnate, with a distinct terminal pinna resembling adjacent lateral pinnae, lower 2-4 pairs of pinnae often deltoid-lanceolate with long basal basiscopic pinnules |
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(20) |
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20 (19) |
Lamina relatively smaller and less dissected when mature, (2 or)3-pinnate, lower pinnae proximally with 1 or 2 pairs of elongated and 1-pinnate pinnules. |
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17 A. simplicior |
+ |
Lamina relatively larger and more dissected, 4-pinnate at base, lower pinnae proximally with 2 or 3 pairs of elongated and 1- or 2-pinnate pinnules. |
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18 A. ziyunshanensis |
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21 (18) |
Lamina 2- or 3-pinnate; stipe bearing brown or dark brown, linear-lanceolate to subulate, denticulate-based, appressed scales above base and rachis throughout |
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(22) |
+ |
Lamina (3 or)4(or 5)-pinnate; stipe bearing brown or coal-black, linear-lanceolate (or wider but often entire-margined) spreading scales above base and rachis throughout, or stipe glabrescent |
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(25) |
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22 (21) |
Lamina caudate-acuminate at apex; lower pinnae with basally elongated pinnules |
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(23) |
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Lamina acuminate to shortly acuminate at apex; pinnae (except basal) often without basally elongated pinnules |
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(24) |
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23 (22) |
Rhizome long creeping; lamina shiny brownish green when dried; sori medial on veinlets. |
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20 A. aristata |
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Rhizome shortly creeping; lamina dull greenish when dried; sori medial or more basal on veinlets. |
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22 A. jinpingensis |
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24 (22) |
Lamina acuminate to attenuate at apex, occasionally abruptly narrowed; lateral pinnae often with basal pinnules longest. |
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21 A. chinensis |
+ |
Lamina acuminate at apex; lateral pinnae with basal pinnules as long as or shorter than adjacent upper ones. |
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23 A. japonica |
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25 (21) |
Lamina subleathery, stiff or firm when dried |
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(26) |
+ |
Lamina thinly herbaceous, soft and tenuous when dried |
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(27) |
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26 (25) |
Lamina thickly papery to subleathery when mature; ultimate segments broader and with aristate or mucronate teeth. |
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24 A. speciosa |
+ |
Lamina papery or thinly papery when mature; ultimate segments often narrower and with only sharply tipped (but non-aristate) teeth. |
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25 A. coniifolia |
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27 (25) |
Stipe above base and rachis throughout bearing sparse, dull brown, relatively thinner scales, or glabrescent. |
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26 A. festina |
+ |
Stipe above base and rachis throughout bearing a number of coal-black and relatively thicker scales. |
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27 A. nigrospinosa |
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28 (16) |
Lamina abruptly caudate at apex and forming a conform linear terminal pinna |
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(29) |
+ |
Lamina toward apex acuminate, attenuate, or abruptly narrowed but acute, or forming a broadly acuminate terminal pinna |
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(31) |
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29 (28) |
Sori marginal or submarginal; indusia margin ciliate at least when young. |
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28 A. amabilis |
+ |
Sori medial; indusia margin entire |
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(30) |
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30 (29) |
Pinnae strongly ascending (forming an angle of less than 30?with rachis), lower 3 or 4 pairs similar in shape and size; ultimate segment margin serrate; indusia with hairs on dorsal surface. |
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30 A. similis |
+ |
Pinnae obliquely spreading (forming an angle of more than 60?with rachis), pinnae gradually reduced toward lamina apex (and sometimes lowest pair of pinnae with long basal basiscopic pinnules); proximal ultimate segments on acroscopic or both margins often shallowly pinnate; indusia glabrous. |
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29 A. hekiana |
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31 (28) |
Lamina 2-pinnate; ultimate segments often broader than 1 cm |
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(32) |
+ |
Lamina 2-pinnate-pinnatifid to usually 3- or 4(or 5)-pinnate; ultimate segments often narrower than 1 cm |
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(35) |
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32 (31) |
Lamina gradually acuminate toward apex |
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(33) |
+ |
Lamina abruptly narrowed toward apex, with a distinct but acuminate terminal pinna |
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(34) |
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33 (32) |
Lamina as long as or shorter than stipe, deltoid-oblong, with a terminal pinna resembling or slightly wider than its lateral ones; ultimate segments not decurrent. |
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31 A. assamica |
+ |
Lamina longer than stipe, often oblong, its 1-pinnate apex much longer and wider than lateral pinnae; ultimate segments decurrent. |
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32 A. yoshinagae |
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34 (32) |
Stipe ochraceous; lamina papery, brownish green when dried; lateral pinnae broader than 5 cm. |
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33 A. pseudoassamica |
+ |
Stipe stramineous; lamina thinly papery, yellowish green when dried; lateral pinnae narrower than 4 cm. |
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34 A. wulingshanensis |
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35 (31) |
Lamina abruptly narrowed toward apex and long acuminate or acute; lowest pinnae linear-lanceolate to lanceolate or basal 1(or 2) pairs of pinnules obviously elongated |
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(36) |
+ |
Lamina acuminate or slightly attenuate at apex; lowest pinnae deltoid or ovate and often much broader, with basal basiscopic pinnules slightly longer |
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(37) |
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36 (35) |
Lamina 2-pinnate-pinnatifid; lateral pinnae linear-lanceolate or lanceolate; ultimate segments often deeply pinnatifid. |
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35 A. hunanensis |
+ |
Lamina 3-pinnate; lower 3(or 4) pinnae deltoid with elongated basal pinnules; ultimate segments (except proximal ones) sharply serrate. |
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36 A. sinorhomboidea |
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37 (35) |
Lamina thickly papery or subleathery, most often 3-pinnate; ultimate segments usually broader than 5 mm |
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(38) |
+ |
Lamina herbaceous or papery, most often 4-pinnate; ultimate segments usually narrower than 5 mm |
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(39) |
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38 (37) |
Pinnules remote; lamina papery and often flexible, shiny yellowish green when dried; veins and costules with brown, appressed articulate hairs abaxially. |
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37 A. nipponica |
+ |
Pinnules contiguous; lamina subleathery and often firm, pale greenish when dried; veins and costules glabrescent abaxially. |
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38 A. ailaoshanensis |
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39 (37) |
Ultimate segments with aristate-tipped teeth; indusia ciliate or erose; lamina firmly herbaceous or papery, often dull pale green when dried. |
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39 A. simulans |
+ |
Ultimate segments with blunt-tipped teeth; indusia entire; lamina thinly herbaceous, often shiny green when dried. |
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40 A. henryi |