1 |
Glomerules all axillary on normal leaf branches; male flowers 4-merous; leaves alternate. |
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1 B. glomerulifera |
+ |
Glomerules all or partly arranged on specialized slender flowering branches; male flowers (3-)4-5-merous; leaves alternate or opposite. |
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(2) |
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2 (1) |
Male glomerules in axils of foliage leaves, female glomerules on specialized long spikelike shoots in terminal and subterminal axils; male flowers 4-5-merous, pedicel 2-5 mm. |
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3 B. zollingeriana |
+ |
Glomerules all arranged on specialized slender flowering shoots, rarely simply axillary; male flowers 4-merous, pedicel very short to absent. |
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(3) |
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3 (2) |
Flowering shoots with tuft of foliage leaves at apex. |
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(4) |
+ |
Flowering shoots without tuft of leaves at apex (except rarely in B. silvestrii). |
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(5) |
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4 (3) |
Leaves not lobed, apex acuminate or caudate. |
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4 B. clidemioides |
+ |
Leaves lobed, apex cuspidate. |
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5 B. umbrosa |
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5 (3) |
Flowering shoots branched; leaves alternate or opposite. |
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(6) |
+ |
Flowering shoots usually unbranched, sometimes loosely branched at base; leaves opposite, seldom alternate in distal part of stem (only in B. allophylla and B. tricuspis). |
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(8) |
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6 (5) |
Flowering shoots in pairs; leaves alternate, abaxially often white tomentose (or strigose in var. tenacissima); achene long stipitate at base. |
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2 B. nivea |
+ |
Flowering shoots solitary; leaves opposite, glabrous or sparsely strigillose abaxially; achene base obtuse or cuneate. |
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(7) |
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7 (6) |
Leaf blade elliptic ovate, apex acuminate; flowering shoots with irregular second order branching; achene base narrowly cuneate. |
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20 B. polystachya |
+ |
Leaf blade ovate-orbicular to suborbicular, apex abruptly caudate; flowering shoots with simple, spikelike branches only; achene base rounded. |
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21 B. conica |
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8 (5) |
Leaf blade often 3-cusped or 3-lobed, rarely 5-lobed at apex. |
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(9) |
+ |
Leaf blade not divided at apex. |
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(12) |
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9 (8) |
Leaf blade often papery, 7-15(-22) cm wide, teeth 10-20 mm. |
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(10) |
+ |
Leaf blade herbaceous, 1-8(-13) cm wide, teeth less than 10 mm. |
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(11) |
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10 (9) |
Leaf blade ovate or broadly ovate, apex sometimes inconspicuously tricuspidate, base often broadly cuneate; spikes or rarely few branched at base. |
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15 B. japonica |
+ |
Leaf blade oblate to 5-angled or oblate to orbicular-ovate, apex often 3-lobed, base often truncate or subcordate; spikes well branched into subpanicle. |
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16 B. tricuspis |
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11 (9) |
Leaves all opposite, apex 3- or 5-cuspidate, base broadly cuneate to subtruncate, teeth 2-10 mm, sparsely strigose or subglabrous on both surfaces. |
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17 B. silvestrii |
+ |
Leaves alternate on upper stems, opposite on lower stems, apex tricuspidate, base broadly cuneate, teeth 1-2.5 mm, long appressed hirsute adaxially, pubescent abaxially. |
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19 B. allophylla |
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12 (8) |
Leaf blade lanceolate or linear-lanceolate. |
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(13) |
+ |
Leaf blade ovate or suborbicular, rarely sublanceolate (in B. hamiltoniana). |
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(16) |
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13 (12) |
Leaf blade herbaceous, lanceolate, smooth adaxially, apex caudate or acuminate; female spikes of usually distant glomerules, often branched at base, straight; achene stipitate or very shortly so. |
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(14) |
+ |
Leaf blade thickly papery, linear-lanceolate, often strongly rugose adaxially, apex tapering to acuminate; female spikes of congested glomerules, simple, nodding; achene long stipitate. |
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(15) |
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14 (13) |
Leaf blade apex entire, margin dentate; flowering shoots unbranched or with a few short irregular branches near base; achene almost without stipe. |
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10 B. formosana |
+ |
Leaf blade apex serrulate, margin finely serrulate; female flowering shoot dichotomously forked at lowermost glomerule; achene stipitate. |
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23 B. ingjiangensis |
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15 (13) |
Leaf blade 14-29 × 2.2-5.5 cm, glabrous or glabrescent adaxially, teeth revolute; female spikes 6-32 cm; perianth tube of female flower 1.6-2.2 mm, apex rounded. |
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24 B. penduliflora |
+ |
Leaf blade 5-24 × 2-6.4 cm, densely appressed strigillose adaxially, teeth flattened; female spikes 4-18 cm; perianth tube of female flower 1-1.5 mm, apex attenuate. |
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25 B. densiflora |
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16 (12) |
Flowering shoots branched at base. |
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(17) |
+ |
Flowering shoots unbranched. |
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(27) |
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17 (16) |
Shrubs. |
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(18) |
+ |
Herbs perennial or subshrubs. |
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(21) |
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18 (17) |
Leaf blade elliptic ovate; flowering shoots with irregular second order branching. |
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20 B. polystachya |
+ |
Leaf blade oblong, narrowly ovate or oblanceolate; flowering shoots with simple, spikelike branches only. |
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(19) |
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19 (18) |
Branches glabrous; spikes very slender, to 26 cm. |
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11 B. hamiltoniana |
+ |
Branches strigose; spikes straight, 4-10 cm. |
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(20) |
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20 (19) |
Branches dark purplish brown; leaf blade oblong or oblong-lanceolate, apex acuminate, tip denticulate; inflorescence unbranched. |
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8 B. lanceolata |
+ |
Branches light brown; leaf blade elliptic, apex acuminate, tip entire; inflorescence with shorter branches from base. |
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9 B. lohuiensis |
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21 (17) |
Leaf margin coarsely 7-14-dentate, teeth gradually larger distally. |
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15 B. japonica |
+ |
Leaf margin dentate, denticulate, or serrulate, all regular. |
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(22) |
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22 (21) |
Leaf blade herbaceous or rarely membranous, suborbicular or orbicular-ovate. |
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(23) |
+ |
Leaf blade usually herbaceous or papery, ovate or oblong. |
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(25) |
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23 (22) |
Leaf base rounded, patent pubescent abaxially. |
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6 B. macrophylla |
+ |
Leaf base truncate or cordate, appressed strigose abaxially. |
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(24) |
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24 (23) |
Leaf blade 6.5-19 × 6-17.5 cm; spikes of distant glomerules to 40 cm. |
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13 B. dolichostachya |
+ |
Leaf blade 5-9.4 × 5-8 cm; spikes of congested glomerules to 5.5 cm. |
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14 B. densiglomerata |
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25 (22) |
Leaf blade papery, oblong, apex denticulate. |
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8 B. lanceolata |
+ |
Leaf blade usually herbaceous, ovate or narrowly ovate, apex entire. |
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(26) |
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26 (25) |
Leaf margin dentate; achene verrucose. |
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10 B. formosana |
+ |
Leaf margin denticulate or serrulate; achene smooth. |
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6 B. macrophylla |
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27 (16) |
Leaf blade ovate-rhombic or rhombic, margin coarsely 3-9-dentate. |
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18 B. spicata |
+ |
Leaf blade ovate or suborbicular, margin serrulate, crenulate, or denticulate. |
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(28) |
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28 (27) |
Leaf blade suborbicular. |
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(29) |
+ |
Leaf blade ovate, rarely also lanceolate (in B. hamiltoniana). |
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(34) |
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29 (28) |
Leaf margin coarsely 9-14-dentate, teeth gradually larger distally. |
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15 B. japonica |
+ |
Leaf margin subregularly dentate, denticulate, or serrulate. |
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(30) |
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30 (29) |
Branchlets densely yellowish tomentose; leaf blade tomentose to densely pubescent on both surfaces. |
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12 B. tomentosa |
+ |
Branchlets strigose; leaf blade strigose adaxially. |
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(31) |
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31 (30) |
Leaf blade patent pubescent abaxially. |
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(32) |
+ |
Leaf blade strigillose abaxially. |
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(33) |
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32 (31) |
Leaf blade sparsely puberulent abaxially. |
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6 B. macrophylla |
+ |
Leaf blade densely pubescent abaxially. |
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13 B. dolichostachya |
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33 (31) |
Herbs perennial to 0.5 m tall; leaf blade 5-9.4 × 5-8 cm; female or bisexual spikes of congested glomerules 2-5 cm. |
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14 B. densiglomerata |
+ |
Subshrubs or shrubs to 3 m tall; leaf blade 6.5-19 × 6-17.5 cm; female spikes of distant glomerules 9-40 cm. |
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13 B. dolichostachya |
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34 (28) |
Branches glabrous; leaf blade narrowly ovate to lanceolate, margin finely crenulate throughout; spikes very slender, to 26 cm. |
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11 B. hamiltoniana |
+ |
Branches strigose; leaf blade ovate or narrowly ovate, margin dentate, denticulate, or serrulate, with entire tip; spikes usually less than 26 cm. |
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(35) |
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35 (34) |
Female spikes 0.8-2 cm; plants 0.4-0.6 m tall. |
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7 B. pilosiuscula |
+ |
Female spikes 4-20 cm; plants 0.7-3.5 m tall. |
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(36) |
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36 (35) |
Shrubs or small trees; petiole to 1.2 cm; female inflorescences often in groups of 2-4; female perianth tube patent-puberulent all over. |
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22 B. siamensis |
+ |
Herbs perennial or subshrubs; petiole to 6 or 13 cm; female inflorescences solitary; female perianth tube strigillose on shoulder. |
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(37) |
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37 (36) |
Leaf teeth 6-20 mm, gradually larger distally. |
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15 B. japonica |
+ |
Leaf teeth 2-5 mm, regular. |
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(38) |
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38 (37) |
Leaf narrowly ovate or oblong-ovate, glabrous or subglabrous adaxially; achene verrucose. |
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10 B. formosana |
+ |
Leaf ovate, often rugose, strigose adaxially; achene smooth. |
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6 B. macrophylla |