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FOC | Family List | FOC Vol. 16 | Boraginaceae

40. Bothriospermum Bunge, Enum. Pl. China Bor. 47. 1833.

斑种草属 ban zhong cao shu

Herbs annual or biennial, hispid and appressed pubescent, hairs discoid at base. Stems erect or prostrate. Leaves alternate, ovate to oblanceolate. Cymes bracteate. Flowers pedicellate. Calyx 5-parted; lobes lanceolate, not enlarged or slightly enlarged in fruit. Corolla rotate, blue or white; tube short; throat appendages 5, scalelike; lobes 5, overlapping in bud, apex obtuse. Filaments extremely short; anthers 5, inserted in corolla tube, included; anthers ovate, obtuse. Ovary 4-parted, valves free, each containing 1 anatropous ovule. Style short, not surpassing valves; stigma capitate. Gynobase flat. Nutlets 4, with cupular emergence adaxially, rounded abaxially, tuberculate, with an oblong, elliptic, or orbicular ringlike aperture; margin of aperture thickened, prominent, entire or sometimes finely dentate; attachment scar basal. Seeds usually not curved.

About five species: Afghanistan, China, India, Indonesia (Java, naturalized), Japan, Kazakhstan, Korea, Kyrgyzstan, Pakistan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Vietnam; all five in China.

1 Nutlets with transverse ringlike aperture on adaxial surface; fruiting calyx strongly accrescent, ca. 8 mm in diam.   1 Bothriospermum chinense
+ Nutlets with longitudinal ringlike aperture on adaxial surface; fruiting calyx slightly accrescent, to 5(-6) mm in diam.   (2)
2 (1) Bracts linear to linear-lanceolate; adaxial aperture of nutlets orbicular   2 Bothriospermum kusnezowii
+ Bracts ovate to oblong; adaxial aperture of nutlets elliptic.   (3)
3 (2) Stems appressed pubescent   5 Bothriospermum zeylanicum
+ Stems spreading hispid and appressed pubescent.   (4)
4 (3) Cymes 3-10 cm; bracts ovate; appendages of corolla throat ca. 0.2 mm   3 Bothriospermum hispidissimum
+ Cymes 10-20 cm; bracts oblong to ovate-lanceolate; appendages of corolla throat ca. 0.8 mm   4 Bothriospermum secundum

Lower Taxa


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