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8. Marattiaceae

合囊蕨科 he nang jue ke

Authors: He Zhaorong & Maarten J. M. Christenhusz

Plants terrestrial (rarely epiphytic, sometimes in streams), evergreen. Rhizomes erect, ascending, or creeping, amylaceous (starchy), polycyclostelic to dictyostelic. Fronds monomorphic and long-lived or dimorphic and fertile ones short-lived (Danaea), simple or compound, 1-4-pinnate or (pedate-)palmate (Christensenia), small (ca. 20 cm) to very large (up to at least 6 m). Stipes fleshy, with a pair of stipulelike appendages at base, often with conspicuous lenticels. Pulvini at base of fronds, at nodes, at bases of pinnae, and in some species at "naked" nodes (without pinnae) along stipe. Veins free, simple or bifurcate, or reticulate (Christensenia), false veins present between true veins or absent. Tissues mucilaginous. Indument of multicellular uniseriate hairs and basifixed or peltate scales. Sori lacking true indusia, usually with paraphyses, superficial, sunken or stalked, elongate along veins, bilateral or radial. Sporangia partially or completely fused into synangia, dehiscing by a slit or pore. Spores trilete or monolete, with bullate, muriform, or stellate ornamentation. Prothalli very large, thalloid, photosynthetic.

Six genera and ca. 100 species: pantropical, Danaea Smith, Eupodium J. Smith, and Marattia Swartz in the Neotropics, Angiopteris, Christensenia, and Ptisana in the Paleotropics; three genera and 30 species (17 endemic) in China.

The delimitation of certain species of Angiopteris is unclear.

The following papers are relevant: Christenhusz (Ann. Univ. Turku., A II, 216: 1-134. 2007); Christenhusz and Toivonen (Biol. Invas. 10: 1215-1228. 2008); Murdock (Amer. J. Bot. 95: 626-641. 2008); and Murdock (Taxon 57: 737-755. 2008).

Ching Ren-chang, Fu Shu-hsia, Wang Chu-hao & Shing Gung-hsia. 1959. Angiopteridaceae and Christenseniaceae. In: Ching Ren-chang, ed., Fl. Reipubl. Popularis Sin. 2: 27-66, 330-345; Zhang Xianchun. 2004. Marattiaceae. In: Zhang Xianchun, ed., Fl. Reipubl. Popularis Sin. 6(3): 246-248.

1 Fronds simple, pedately lobed to palmately compound; venation reticulate; synangium radial.   2 Christensenia
+ Fronds pinnate (sometimes simple when plants juvenile); venation free, simple or bifurcate; synangium ovate   (2)
2 (1) Sporangia fully fused into synangia, synangia bivalved.   1 Ptisana
+ Sporangia fused basally but free apically, synangia composed of globose sporangia.   3 Angiopteris

Lower Taxa


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