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FOC | Family List | FOC Vol. 11 | Euphorbiaceae

3. Cleistanthus J. D. Hooker ex Planchon, Icon. Pl. 8: t. 779. 1848.

闭花木属 bi hua mu shu

Authors: Bingtao Li & Stefan Dressler

Leiopyxis Miquel; Paracleisthus Gagnepain.

Trees or shrubs, monoecious or dioecious; indumentum of simple hairs. Leaves alternate, petiolate; stipules persistent or caducous; leaf blade simple, entire, venation pinnate. Flowers in axillary glomerules or spikes, small, sessile or sometimes shortly pedicellate. Male flowers: sepals 4-6, valvate; petals 4-6, small, scalelike; disk cupular or cushionlike; stamens 5; filaments connate up to half their length, topped with rudimentary ovary; anthers usually ovoid, dorsifixed, extrorse, anther bilocular, longitudinally dehiscent. Female flowers: sepals and petals as in male; disk cup-shaped or conical, surrounding basal ovary, sometimes as long as ovary, tearing into lobes during fruit development, rarely absent; ovary 3(or 4)-celled; ovules 2 in each cell; styles 3, bifid at apex. Fruit a capsule, subglobose, (bi- or) trivalvate when mature, sometimes stipitate (carpopodium), epicarp thin, endocarp bony, inside fibrous; columella and sepals persistent. Seeds (1 or)2 in each cell; cotyledons broad, thinly to thickly fleshy; endosperm copious to submembranous. x = 11.

About 141 species: tropical Africa, Asia, and Australia; seven species in China.

1 Ovary and capsules pubescent or hirsute   (2)
+ Ovary and capsules glabrous   (4)
2 (1) Branchlets and petioles rusty tomentose; leaves pubescent abaxially.   1 C. tomentosus
+ Branchlets (except C. sumatranus young branchlets puberulent), petioles, and leaves abaxially glabrous   (3)
3 (2) Leaf blade 3-10(-15) × 2-5(-7) cm, apex caudate-acuminate; lateral veins usually obscure; bract margins glabrous; petal apex entire.   2 C. sumatranus
+ Leaf blade 14-30 × 8-15 cm, apex obtuse; lateral veins prominent; bracts ciliate at margin; petal apex crenulate.   3 C. macrophyllus
4 (1) Female disk as long as or longer than ovary   (5)
+ Female disk much shorter than ovary, only to base of ovary   (6)
5 (4) Leaf blade apex long acuminate; male pedicels 5-10(-13) mm; female disk apex pubescent.   4 C. pedicellatus
+ Leaf blade apex shortly acuminate; male flowers sessile; female disk apex glabrous.   6 C. petelotii
6 (4) Leaf blade apex long acuminate, lateral veins 8-10 pairs; disk margin entire.   5 C. tonkinensis
+ Leaf blade apex obtuse or retuse, lateral veins 7-9 pairs; disk margin erose or with 5 crenulations.   7 C. concinnus

Lower Taxa


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