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10. Cleyera Thunberg, Nov. Gen. Pl. 68. 1783.
红淡比属 hong dan bi shu
Sakakia Nakai; Tristylium Turczaninow.
Shrubs or trees, evergreen. Leaves petiolate; leaf blade margin entire or serrulate. Flowers bisexual, axillary, solitary or several in a cluster. Pedicel 1 cm or more, apically thickened; bracteoles 2, caducous, small, inserted near pedicel apex. Sepals 5, imbricate, unequal, basally slightly connate; outer sepals persistent, small. Petals 5, imbricate, basally connate. Stamens 25-30; filaments distinct, glabrous; anthers basifixed, 2-loculed, longitudinally dehiscent, with filiform trichomes, connective apiculate. Ovary usually glabrous, 2- or 3-loculed with 8-16 ovules per locule, placentation axile; style 1, persistent, slender, elongated, apically 2- or 3-lobed. Fruit baccate, ovoid to oblate, with several seeds per locule. Seeds blackish brown, reniform-globose to compressed globose, foveolate, shiny, glabrous; endosperm sparse; embryo curved.
About 24 species: China, N India, Japan, Korea, Myanmar, Nepal, N Vietnam; tropical America; nine species (seven endemic) in China.
Unlike Cleyera japonica, the Thunberg description of the genus Cleyera did not include elements of a mixed collection with Ternstroemia.
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| 1 |
Fruit long ovoid, oblong, obconic, or ovoid-ellipsoid |
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(2) |
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Fruit globose |
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(4) |
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| 2 (1) |
Leaf blade ovate to obovate-oblong, margin entire; sepals orbicular. |
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8 C. obovata |
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Leaf blade oblong-elliptic or rarely oblong-oblanceolate, margin serrulate, sparsely serrate, or rarely subentire; sepals suborbicular, triangular, or ovate-triangular |
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(3) |
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| 3 (2) |
Leaf blade abaxially reddish brown glandular punctate; sepals suborbicular. |
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7 C. obscurinervia |
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Leaf blade abaxially not punctate; sepals triangular to ovate-triangular. |
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9 C. longicarpa |
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| 4 (1) |
Leaf blade abaxially not punctate |
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(5) |
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Leaf blade abaxially densely or sparsely reddish or reddish brown glandular punctate |
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(7) |
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| 5 (4) |
Sepals ovate-triangular, apex acute with a black mucro; petiole 3-5 mm; pedicel 0.5-0.8 cm. |
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3 C. parvifolia |
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Sepals ovate to orbicular, apex rounded and without a mucro; petiole 7-10(-12) mm; pedicel 1-2(-3) cm |
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(6) |
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| 6 (5) |
Leaf blade margin entire, midvein adaxially slightly impressed. |
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1 C. japonica |
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Leaf blade margin conspicuously serrate, crenate serrate, or sparsely serrate, midvein adaxially raised. |
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2 C. lipingensis |
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| 7 (4) |
Leaf blade oblong, secondary veins 20-28 on each side of midvein; sepals oblong to ovate-oblong, thick. |
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4 C. pachyphylla |
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Leaf blade elliptic, oblong-elliptic, narrowly elliptic, or oblanceolate-elliptic, secondary veins 8-12 on each side of midvein; sepals ovate to orbicular, thin |
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(8) |
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| 8 (7) |
Leaf blade elliptic to oblong-elliptic, margin entire, midvein adaxially raised, secondary veins 12-14 on each side of midvein. |
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5 C. yangchunensis |
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Leaf blade narrowly elliptic to oblanceolate-elliptic, margin serrulate to sparsely serrate, midvein adaxially slightly impressed, secondary veins 8-12 on each side of midvein. |
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6 C. incornuta |
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List of lower taxa
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