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FOC | Family List | FOC Vol. 16 | Gentianaceae

18. Comastoma (Wettstein) Toyokuni, Bot. Mag. (Tokyo). 74: 198. 1961.

喉花草属 hou mao hua shu

Gentiana sect. Comastoma Wettstein, Oesterr. Bot. Z. 46: 174. 1896.

Herbs annual with fibrous rootlets or perennial with short rhizomes. Stems branched from base (simple in C. disepalum). Inflorescences racemelike, sometimes reduced to a solitary terminal flower. Flowers 4 or 5merous. Calyx lobed nearly to base, without an intracalycular membrane. Corolla tubular, tubular-funnelform, or salverform; lobes each with 1 or 2 basal scales; scales fringed with nonvascularized white fimbriae; plicae absent. Nectaries at base of corolla tube. Stamens inserted on corolla tube; filaments white. Capsules 2valved, many seeded. Seed coat smooth.

Fifteen species: North America, Asia, Europe; 11 species in China.


1 Calyx 2lobed, enveloping corolla tube   1 Comastoma disepalum
+ Calyx 4 or 5lobed, not enveloping corolla tube.   (2)
       
2 (1) Perennials.   (3)
+ Annuals or biennials.   (4)
       
3 (2) Leaves mostly restricted to near base of stem; stem leaves few pairs; corolla 1.4-2.5 cm, lobes each bearing 2 scales   2 Comastoma cyananthiflorum
+ Basal leaves absent or poorly developed; stem leaves 4-7 pairs; corolla 0.6-1.3 cm, lobes each bearing 1 scale   3 Comastoma stellariifolium
       
4 (2) Corolla salverform.   (5)
+ Corolla tubular.   (7)
       
5 (4) Calyx ca. 1/2 as long as corolla, lobes falcate, apex acuminate; stem leaf blades 3-4(-6) mm wide   6 Comastoma falcatum
+ Calyx 1/4-1/3 as long as corolla, lobes erect, apex acute to obtuse; stem leaf blades 5-13 mm wide.   (6)
       
6 (5) Stem leaf blades oblong to broadly ovate; calyx lobes ovate-lanceolate to ovate; filaments pilose at base   4 Comastoma traillianum
+ Stem leaf blades lanceolate to elliptic; calyx lobes linear-lanceolate; filaments glabrous   5 Comastoma muliense
       
7 (4) Stem leaf blade bases obtuse to rounded, subamplexicaul; corolla lobes shorter than tube, suberect   7 Comastoma pulmonarium
+ Stem leaf blade bases narrowed or cuneate (rounded and abruptly contracted in C. henryi), not amplexicaul; corolla lobes equaling tube, spreading (erect in C. henryi).   (8)
       
8 (7) Stem leaves petiolate; leaf blades widest above middle; calyx lobes oblong-spatulate to narrowly elliptic   8 Comastoma henryi
+ Stem leaves sessile; leaf blades oblong or widest at or below middle; calyx lobes lanceolate to ovate.   (9)
       
9 (8) Corolla dark blue to blue-purple with dark blue veins and pale yellow base   11 Comastoma pedunculatum
+ Corolla uniformly blue or pale blue.   (10)
       
10 (9) Calyx lobes acuminate, margin undulate rugose and revolute   9 Comastoma polycladum
+ Calyx lobes apiculate, margin flat and smooth   10 Comastoma tenellum

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