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FOC | Family List | FOC Vol. 2-3 | Cystopteridaceae

4. Cystopteris Bernhardi in Schrader, Neues J. Bot. 1(2): 26. 1805.

冷蕨属 leng jue shu

Authors: Zhongren Wang & Christopher Haufler

Cyste Dulac; Cystea J. Smith; Filix Adanson (1763), not Séguier (1754), nor Ludwig (1757); Rhizomatopteris A. P. Khokhrjakov.

Plants small, summer-green. Rhizomes long or shortly creeping, blackish brown, glabrate or densely red-brown pubescent, sparsely scaly; scales brown or brownish, thin, ovate to broadly lanceolate. Fronds distant, approximate, or caespitose, thin; stipe dark brown at base, upper part stramineous or chestnut-colored; lamina 2- or 3-pinnate, rarely 4-pinnate or more divided, ovate-lanceolate, ovate-triangular, or subpentagonal; pinnae with anadromous pinnules, shortly stalked, unequally sided or subequilateral at base; minutely toothed at segment margin; veins free, forked or pinnate, terminating in teeth or emarginations. Lamina thinly herbaceous or herbaceous, green when dried, stipe, rachis, costae, and costules with few small scales at base, and multicellular hairs or unicellular glandular hairs, lamina usually glabrate. Sori orbicular, abaxial on veins, sorophore slightly protuberant, annulus of sporangium consisting of 14-16 cells; indusia ovate, lanceolate, orbicular, or shallowly cup-shaped, attached at proximal side of sorophore, membranous, persistent, covering sporangia when young, hidden at maturity. Spores bilateral, dark brown, bean-shaped, perispore usually with spines, rarely with folds. x = 42.

More than 20 species and many hybrids: mainly in the temperate and frigid-temperate zones and in tropical mountains; 11 species (six endemic) in China.

The following taxon is excluded from the present treatment, pending further research: Cystopteris maborasensis Masamune (Trans. Nat. Hist. Soc. Formosa 28: 141. 1938), described from Taiwan.

1 Rhizomes shortly creeping or ascending; fronds approximate or caespitose; lamina lanceolate to broadly lanceolate; veins terminating in teeth, rarely in emarginations   (2)
+ Rhizomes long creeping with fronds distant; lamina broadly ovate, ovate-triangular, or subpentagonal; veins terminating in emarginations   (5)
2 (1) Veins terminating in teeth   (3)
+ Veins terminating in emarginations   (4)
3 (2) Perispore echinate; on calcareous soil.   1 C. fragilis
+ Perispore rugose or verrucose, not echinate; on acidic soil.   2 C. dickieana
4 (2) Plants usually less than 10 cm tall; stipe slender, slightly longer than lamina or nearly as long as lamina, chestnut-colored.   3 C. kansuana
+ Plants up to 29 cm tall; stipe thick (ca. 1 mm), shorter than lamina, dark purple at base.   4 C. guizhouensis
5 (1) Lamina subpentagonal with basal basiscopic pinnules of basal pinnae prominently elongated   (6)
+ Lamina broadly ovate or ovate-triangular with basal basiscopic pinnules of basal pinnae not elongated   (7)
6 (5) Ultimate segments not revolute at margin; spore surface shortly spinose or verrucose.   5 C. montana
+ Ultimate segments revolute at margin; spore surface rugately folded.   6 C. modesta
7 (5) Indusia glandular; plants slender.   7 C. sudetica
+ Indusia not glandular, plants moderately stout   (8)
8 (7) Lamina with short glandular hairs.   8 C. tibetica
+ Lamina lacking short glandular hairs   (9)
9 (8) Fronds up to 50 cm or longer; lamina to 4-pinnatisect; perispore densely echinate.   9 C. deqinensis
+ Fronds less than 50 cm; lamina 2- or 3-pinnate; perispore sparsely echinate   (10)
10 (9) Ultimate segments 3-5 mm wide; indusia attached to ca. 1/2 of circumference of receptacle.   10 C. moupinensis
+ Ultimate pinnules or segments 5-7 mm wide; indusia attached to ca. 2/3 of circumference of receptacle.   11 C. pellucida

  • List of lower taxa


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