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FOC | Family List | FOC Vol. 2-3 | Davalliaceae

3. Davallia Smith, Mém. Acad. Roy. Sci. (Turin). 5: 414. 1793.

骨碎补属 gu sui bu shu

Parestia C. Presl; Stenolobus C. Presl; Trogostolon Copeland; Wibelia Bernhardi (Oct-Dec 1801), not G. Gaertner, B. Meyer & Scherbius (1801).

Plants moderate-sized, epiphytic or sometimes terrestrial. Rhizome long creeping, with dark, peltate scales. Fronds remote, long stipitate, monomorphic or dimorphic; stipe articulate to short phyllopodia, terete or slightly winged; lamina bipinnate to 4-pinnate-pinnatifid, deltoid or pentagonal, firmly leathery or sometimes thickly herbaceous, glabrous; ultimate segments with crenate or lobed margins. Veins free, usually forked, terminating in lobes or crenations of cartilaginous margin. Sori terminal on veins, on small oblique lobes or crenations; indusium extrorse, elongate toward margin, attached at base and sides, long stalked; annulus longitudinal, consisting of ca. 14 thickened cells; spores monolete, elliptic, smooth, translucent; perispore absent. n = 10, (40).

About 40 species: from Atlantic Ocean islands through Africa and S Asia to Malaysia, Japan, NE Australia, and Pacific islands; six species (one endemic) in China.

The following taxa are excluded from the present treatment, pending further research: Davallia triangularis Baker (Ann. Bot. 5: 202. 1891), described from Yunnan, and D. lorrainii Hance (Ann. Sci. Nat., Bot., sér. 5, 5: 254. 1866; D. trichomanoides Blume var. lorrainii (Hance) Holttum) recorded from Yunnan by W. M. Chu & Z. R. He (Acta Bot. Yunnan. 22: 261. 2000).

1 False veins present.   1 D. denticulata
+ False veins absent   (2)
2 (1) Fronds ca. 1 m tall; lamina 55-100 ?40-90 cm.   2 D. divaricata
+ Fronds ca. 0.5 m tall; lamina 8-35(-90) ?8-30(-40) cm   (3)
3 (2) Lamina papery; rhizome 4-5 mm in diam., scales gray-brown, shaggy.   3 D. trichomanoides
+ Lamina leathery or thickly papery; rhizome 8-10 mm in diam., scales dark brown and overlapping   (4)
4 (3) Rhizome scales very dark brown; lamina thickly papery; apex of pinnae obtuse or acute.   4 D. sinensis
+ Rhizome scales red-brown; lamina thickly leathery; apex of pinnae acuminate   (5)
5 (4) Rhizome scales ovate-lanceolate, with a long subulate apex and ciliate margin; indusia attached along sides, tubular.   5 D. solida
+ Rhizome scales triangular, with an attenuate to obtuse apex and glabrous margin; indusia basifixed, broadly crateriform.   6 D. napoensis

Lower Taxa


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