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FOC | Family List | FOC Vol. 15 | Myrsinaceae

4. Embelia N. L. Burman, Fl. Ind. 62. pl. 23. 1768.

酸藤子属 suan teng zi shu

Shrubs, becoming lianous at maturity, or lianas. Leaves alternate, distichous, or pseudoverticillate. Inflorescences racemose, paniculate, umbellate or corymbose, bracteate at base. Pedicel subtended by one persistent floral bract. Flowers 4- or 5-merous, bisexual or functionally unisexual and plants dioecious, polygamous, or monoecious. Sepals united at base. Petals free or united at base, glandular granulose inside and often along margin. Stamens adnate to base of petals, rarely free, normally exserted in staminate flowers, reduced in pistillate ones; filaments free; anthers dehiscing longitudinally, connectives usually punctate abaxially. Pistillode reduced and style included in staminate flowers; ovary globose or ovoid; ovules ca. 4, uniseriate. Style exserted; stigma discoid or capitate, sometimes slightly lobed. Fruit drupes, prominently punctate, 1-seeded; endocarp crusty or rarely bony. Seeds subglobose, covered by membranous remnants of placenta; endosperm ruminate; embryo terete, transverse.

About 140 species: Africa, SE Asia, Australia, Pacific Islands; 14 species in China.

1 Inflorescences pendent, corymbose umbellate.   (2)
+ Inflorescences erect, racemose or paniculate.   (7)
2 (1) Leaf margin entire.   (3)
+ Leaf margin sharply serrate or crenate, with spiny or acute to obtuse teeth.   (4)
3 (2) Leaf blade oblanceolate or obovate, base cuneate; petiole 5-7 mm   5 Embelia laeta
+ Leaf blade broadly ovate or ovate, base truncate or subcordate; petiole 0.5-1 mm   6 Embelia parviflora
4 (2) Leaf apex mucronate, obtuse or acute.   (5)
+ Leaf apex not mucronate, caudate to acuminate.   (6)
5 (4) Leaf blade glossy, base cordate; petiole subterete, (2-)2.5-3.5(-5) mm   1 Embelia polypodioides
+ Leaf blade dull, base obtuse or acute; petiole deeply canaliculate, 4-5 mm   2 Embelia procumbens
6 (4) Branchlets glandular papillose-tomentose; leaf blade teeth spinose; perianth leathery; sepals broadly ovate, apex acute or obtuse; petals involute, margin glabrous   3 Embelia pauciflora
+ Branchlets glandular puberulent; leaf blade teeth obtuse to acute; perianth papery; sepals narrowly ovate, apex acuminate or attenuate; petals flat, margin glandular granulose   4 Embelia henryi
7 (1) Inflorescences paniculate; petiole marginate.   (8)
+ Inflorescences racemose; petiole canaliculate.   (10)
8 (7) Branchlets and inflorescence rachis reddish glandular papillose-puberulent, early glabrescent; sepals prominently black punctate, margin glabrous, acute to obtuse; petals prominently pellucid punctate-lineate, margin glandular granulose   9 Embelia floribunda
+ Branchlets and inflorescence rachis tawny papillose-tomentose; perianth inconspicuously pellucid punctate, margin long glandular papillose; sepals acuminate.   (9)
9 (8) Inflorescence branches spicate; leaf margin prominently black punctate   7 Embelia sessiliflora
+ Inflorescence branches racemose; leaf margin punctate   8 Embelia ribes
10 (7) Branchlets 5-7 mm in diam.; leaf blade leathery to cartilaginous, margin entire.   (11)
+ Branchlets 2-4.5 mm in diam.; leaf blade papery to subleathery, margin serrate or subentire.   (12)
11 (10) Branchlets and leaf midrib rusty stellate tomentose; abaxial leaf surface with numerous small reddish scales, prominently red and black punctate   13 Embelia gamblei
+ Branchlets and leaf midrib glabrous; abaxial leaf surface glabrous, sparsely and inconspicuously black punctate   14 Embelia carnosisperma
12 (10) Leaf blade glossy, margin regular, serrate or serrulate throughout; sepals erose, inconspicuously pellucid or brown punctate, apex obtuse to acutish   10 Embelia vestita
+ Leaf blade dull, margin irregular, subentire or roughly serrulate-dentate distally; sepals entire, prominently black punctate, apex acuminate.   (13)
13 (12) Branchlets glabrous; leaf base cuneate; fruit longitudinally ribbed, 1-1.5 cm in diam.   11 Embelia undulata
+ Branchlets minutely reddish papillate-puberulent; leaf base obtuse to rounded; fruit smooth, 0.3-0.5 cm in diam.   12 Embelia scandens

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