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FOC | Family List | FOC Vol. 17 | Lamiaceae

28. Eremostachys Bunge, Fl. Altaic. 2: 414. 1830.

沙穗属 sha sui shu

Herbs perennial, erect. Basal leaves largest, margin coarsely serrate to incised-pinnatipartite. Verticillasters many flowered, widely spaced or in long robust spikes, densely lanate-villous or glabrous. Flowers sessile. Calyx tubular-campanulate to broadly funnelform, 5-toothed, sometimes limb dilated, 5-apiculate; teeth short, broadly truncate, rounded, ovate or triangular, apex spiny; sinuses between teeth with ovate-triangular appendages. Corolla 2-lipped; tube mostly included, narrow; upper lip narrow, galeate or falcate, concave, narrowed at base, bearded or villous inside and on margin; lower lip spreading, 3-lobed, middle lobe largest. Stamens 4, anterior 2 longer; some filaments with comblike-fimbriate appendages at base; anther cells 2, divaricate. Style unequally 2-cleft, anterior lobe larger. Nutlets obovoid, triquetrous, subtruncate, densely bearded at apex.

About 60 species (or only five in the strict sense): Kazakhstan, Kyrgyzstan, Mongolia, Russia, Tajikistan, Turkmenistan; SW Asia; five species in China.

Several botanists have recently rearranged the generic placings of Eremostachys and Phlomis (cf. Adylov, Kamelin, & Makhmedov, Novosti Sist. Vyssh. Rast. 23: 110-114, 1986; Vvedensky, Conspect. Fl. As. Med. 9: 74-113, 1987). By adopting their concepts species 1 and 2 would be placed in Phlomoides along with all but one species of Phlomis recorded from China, species 3 and 4 in Paraeremostachys and only E. moluccelloides would remain in Eremostachys.

1 Calyx funnelform, dilated at apex, especially in fruit, membranous   5 Eremostachys moluccelloides
+ Calyx tubular or tubular-campanulate, not dilated at apex, not membranous.   (2)
2 (1) Basal leaves pinnatipartite to bipinnatipartite   1 Eremostachys speciosa var. viridifolia
+ Basal leaves not pinnatipartite, irregularly shallow serrate or crenate.   (3)
3 (2) Bracts leaflike, separated at base; leaves irregularly shallow serrate   2 Eremostachys fulgens
+ Bracts spinescent, united at base in 3s; leaves crenate.   (4)
4 (3) Plants sparsely white hairy; calyx teeth with long soft spines   3 Eremostachys phlomoides
+ Plants densely white hairy; calyx teeth with robust rigid spines   4 Eremostachys desertorum

Lower Taxa


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