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FOC | Family List | FOC Vol. 15 | Oleaceae

2. Fraxinus Linnaeus, Sp. Pl. 2: 1057. 1753.

衿属 qin shu

Wei Zhi; Peter S. Green

Trees or rarely shrubs, deciduous or rarely evergreen. Leaves odd-pinnate, opposite or rarely whorled at branch apices; petiole and petiolule often basally thickened. Inflorescences terminal or axillary toward end of branches, or lateral on branches of previous year, paniculate; bracts linear to lanceolate, caducous or absent. Flowers small, unisexual, bisexual, or polygamous. Calyx 4-toothed or irregularly lobed, sometimes absent. Corolla white to yellowish, 4-lobed, divided to base or absent. Stamens 2, inserted at base of corolla lobes; filaments short, exserted at anthesis. Ovules 2 in each locule, pendulous. Style short; stigma ± 2-cleft. Fruit a samara with apically elongated wing. Seeds usually 1, ovate-oblong; endosperm fleshy; radicle erect.

About 60 species: mostly in temperate regions and subtropics of the Northern Hemisphere; 22 species in China.

Some species are widely used in China for commercial timber and the bark for medicinal drugs.


1 Panicles from lateral leafless buds of previous year; flowers appear before leaves.   (2)
+ Panicles terminal or from lateral buds of this year; flowers appearing with or after leaves.   (6)
       
2 (1) Leaflet blade 1.7-5(-5.5) × 0.5-1.5 cm; panicles congested, 0.5-1.5 cm.   (3)
+ Leaflet blade (2.5-)5-14(-20) × (1-)2-5 cm; panicles lax, 5-20 cm.   (4)
       
3 (2) Leaflets crenate distally, entire proximally; 4 primary veins on each side of midrib   18 Fraxinus xanthoxyloides
+ Leaflets sharply serrate; 6 or 7 primary veins on each side of midrib   19 Fraxinus hupehensis
       
4 (2) Flowers with calyx; samara not twisted   20 Fraxinus platypoda
+ Flowers without calyx; samara conspicuously twisted.   (5)
       
5 (4) Branchlets nearly 4-angled; leaves opposite; leaflets subsessile; inflorescences 15-20 cm, spreading   21 Fraxinus mandschurica
+ Branchlets terete; leaves in whorls of 3 at top of branches; leaflets petiolulate; inflorescences ca. 5 cm   22 Fraxinus sogdiana
       
6 (1) Flowers without corolla.   (7)
+ Flowers with corolla.   (8)
       
7 (6) Leaflets broadly ovate, ovate, to lanceolate, 2-7 cm wide   16 Fraxinus chinensis
+ Leaflets narrowly lanceolate, 1-1.8(-2.2) cm wide   17 Fraxinus baroniana
       
8 (6) Bracts persistent; winter buds naked.   (9)
+ Bracts caducous or absent; winter buds scaly or tomentose.   (14)
       
9 (8) Leaflets 3(-5), serrate to crenate; calyx teeth deltate.   (10)
+ Leaflets 5 or more, entire; calyx lobes truncate or broadly deltate.   (11)
       
10 (9) Leaflets regularly serrate, abaxially tomentose   5 Fraxinus trifoliolata
+ Leaflets remotely crenate, abaxially glabrous   6 Fraxinus punctata
       
11 (9) Leaflets 5-7(-11), petioule 5-10 mm, usually glabrous on both surfaces   2 Fraxinus griffithii
+ Leaflets 9-25, petioule 0-5 mm, midrib at least abaxially hairy.   (12)
       
12 (11) Leaf rachis rusty tomentose; leaflets scurfy hairy or glabrescent   1 Fraxinus ferruginea
+ Leaf rachis and adaxial surface of leaflets densely brown tomentose, leaflets sometimes glabrescent.   (13)
       
13 (12) Leaflets 9-15, densely tomentose adaxially   3 Fraxinus malacophylla
+ Leaflets (11-)15-25, tomentose or glabrescent   4 Fraxinus retusifoliolata
       
14 (8) Petiolules 5-15 mm.   (15)
+ Petiolules 0-5 mm.   (17)
       
15 (14) Leaflets 2-5 cm, margin deeply serrate to incised   15 Fraxinus bungeana
+ Leaflets 8-15 cm, margin serrate or serrulate.   (16)
       
16 (15) Leaflets 7 or 9, abaxially hairy; samara densely red scurfy hairy   10 Fraxinus floribunda
+ Leaflets 3-5(-7), glabrous; samara not as above   11 Fraxinus insularis
       
17 (14) Leaflets 7 or 9.   (18)
+ Leaflets 3-5(-7).   (19)
       
18 (17) Leaflets lanceolate, abaxially densely felty; leaf axis subterete   7 Fraxinus sikkimensis
+ Leaflets ovate-oblong, glabrous or villous on veins abaxially; leaf axis grooved   9 Fraxinus paxiana
       
19 (17) Leaflets 2-5 cm, broadly ovate, rhomboid to ovate or broadly lanceolate to broadly elliptic, with 4-6 primary veins on each side of midrib, margin deeply serrate to incised   15 Fraxinus bungeana
+ Leaflets 3-12 cm, elliptic, lanceolate, or ovate-lanceolate to broadly lanceolate, with 6-12 veins on each side of midrib, margin coarsely serrate, serrate, or entire.   (20)
       
20 (19) Young shoots puberulent; petiolules pubescent or sparsely crisped hairy.   (21)
+ Young shoots glabrous; petiolules glabrous.   (22)
       
21 (20) Petiolules 8-15 mm, sparsely crisped hairy; leaflets with 10-12 primary veins on each side of midrib, margin irregularly and coarsely serrate   8 Fraxinus depauperata
+ Petiolules 0-5 mm, pubescent; leaflets with 7-10 primary veins on each side of midrib, margin entire or serrulate in upper part   14 Fraxinus sieboldiana
       
22 (20) Leaflet margin serrate, blade sparsely dotted abaxially   12 Fraxinus odontocalyx
+ Leaflet margin serrulate, blade not dotted abaxially   13 Fraxinus stylosa

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