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FOC | Family List | FOC Vol. 19 | Annonaceae

8. Goniothalamus (Blume) J. D. Hooker & Thomson, Fl. Ind. 1: 105. 1855.

哥纳香属 ge na xiang shu

Authors: Bingtao Li & Michael G. Gilbert

Polyalthia sect. Goniothalamus Blume, Fl. Javae, Annonaceae, 71, 79. 1830.

Trees or erect shrubs. Petiole short; leaf blade large, secondary veins arcuately looped near margin. Inflorescences axillary or extra-axillary, 1-flowered or several flowers in fascicles. Pedicel bracteolate at base. Sepals 3, valvate. Petals 6, in 2 whorls, with each whorl valvate; outer petals flat, thick; inner petals small, base shortly clawed, apex forming a cap. Stamens many; anther locules linear or oblong, extrorse; connectives oblong or clavate, 3-angular, apex truncate or rarely rounded. Carpels many; ovules 1-10 per carpel, ± basal; styles elongate; stigmas entire or apex 2-lobed. Fruit apocarpous; monocarps subsessile to shortly stipitate, oblong-ellipsoid or ovoid. Seeds 1-10 per monocarp.

About 130-140 species: tropical and subtropical Asia; 11 species (five endemic) in China.

A record of Goniothalamus macrophyllus (Blume) J. D. Hooker & Thomson from the Flora area (X. L. Hou, http://www.abclunwen.com/ lunwen-free-387606/; accessed on 25 Nov 2010) was based on a misidentification of Polyalthia simiarum.


1 Leaf blade margin and midvein abaxially densely rust-colored hirsute   (2)
+ Leaf blade glabrous throughout   (3)
       
2 (1) Leaf blade 56-76 × 13-19 cm; monocarps 6-9 × ca. 2 cm.   1 G. cheliensis
+ Leaf blade 20-41 × 5.5-11.5 cm; monocarps 2-3 × 0.6-0.8 cm.   2 G. donnaiensis
       
3 (1) Inflorescences 3- or 4-flowered; leaf blade abaxially grayish green; inner petals ca. 2 × as long as outer petals.   11 G. laoticus
+ Inflorescences 1- or 2-flowered; leaf blade abaxially green, sometimes drying brownish; inner petals much shorter than outer petals   (4)
       
4 (3) Leaf blade narrowly lanceolate to oblong-lanceolate   (5)
+ Leaf blade oblong, oblong-elliptic, elliptic, or obovate   (7)
       
5 (4) Branchlets and petioles pubescent; stigmas deeply 2-cleft.   3 G. chinensis
+ Branchlets and petioles glabrous; stigmas entire or shallowly 2-cleft   (6)
       
6 (5) Ovaries pubescent; ovules 2 per carpel.   4 G. gardneri
+ Ovaries glabrous; ovules 1 per carpel.   5 G. gabriacianus
       
7 (4) Leaf blade thinly papery, secondary veins adaxially flat to depressed; inflorescences 2-flowered   (8)
+ Leaf blade papery to leathery, secondary veins adaxially raised; inflorescences 1-flowered   (9)
       
8 (7) Pedicel ca. 4 mm; styles ca. 2 × as long as ovaries; ovules basal; leaf blade secondary veins 7-9 on each side of midvein.   6 G. yunnanensis
+ Pedicel 9-15 mm; styles 1-1.5 × as long as ovaries; ovules lateral; leaf blade secondary veins 9-21 on each side of midvein.   7 G. leiocarpus
       
9 (7) Petioles and monocarps pubescent to puberulent.   8 G. amuyon
+ Petioles and monocarps glabrous   (10)
       
10 (9) Flowers to 7 cm, ca. 5.5 cm in diam.; outer petals oblong-lanceolate; ovules 2 per carpel; monocarps ca. 1.5 × 1.3 cm.   9 G. calvicarpus
+ Flowers to 2.5 cm, ca. 2.5 cm in diam.; outer petals broadly ovate; ovules 6 per carpel; monocarps 3-6 × 2-2.5 cm.   10 G. howii

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