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FOC | Family List | FOC Vol. 16 | Asclepiadaceae

25. Hoya R. Brown, Prodr. 459. 1810.

球兰属 qiu lan shu

Centrostemma Decaisne; Otostemma Blume.

Subshrubs or lianas, epiphytic or epilithic, often twining or climbing by adventitious roots. Leaves opposite, fleshy, papery, or membranous. Inflorescences extra-axillary or sometimes terminal, umbel-like, producing a succession of globose or flat-top-ped clusters; old rachis thick, with contiguous pedicel scars. Calyx small, with basal glands. Corolla fleshy, rotate, reflexed, [or campanulate]; lobes valvate, often densely hairy or scurfy inside. Corona lobes 5, depressed patent or adnate vertically to gynostegium, fleshy, glossy when dried, inner angle a tooth incumbent on anthers, outer angle rounded or acute; margin revolute, longitudinally grooved basally. Stamens short connate, apical membrane of anthers appressed against stigma head; pollinia 2 per pollinarium, oblong, erect, with raised, translucent margin. Stigma head discoid, rounded or subapiculate. Follicles often solitary, cylindric-fusiform.

At least 100 species: SE Asia to Oceania; 32 species in China.

1 Leaves 3-veined from base   2 Hoya pottsii
+ Leaves pinnately veined, or veins obscure.   (2)
2 (1) Inflorescences 4- or 5-flowered, terminal; leaves 1-1.5 cm   32 Hoya chinghungensis
+ Inflorescences many flowered, usually extra-axillary, rarely terminal (H. lii); leaves more than 2 cm.   (3)
3 (2) Outer angle of corona lobes rounded.   (4)
+ Outer angle of corona lobes acute.   (13)
4 (3) Leaves villous or pubescent.   (5)
+ Leaves glabrous.   (6)
5 (4) Leaf blade broadly oblong or long quadrate, villous, apex mucronate; corolla lobes as long as tube   23 Hoya villosa
+ Leaf blade elliptic, oblong, elliptic-lanceolate, or rarely suborbicular, pubescent, apex short acuminate to obtuse; corolla lobes longer than tube   24 Hoya lyi
6 (4) Leaf blade orbicular to widest above middle, 3-6 cm wide.   (7)
+ Leaf blade of other shapes, if more than 3 cm wide then widest below middle.   (9)
7 (6) Leaf blade spatulate to oblanceolate, 10-21 cm, lateral veins 10 or more pairs; flower clusters flat topped; corolla white with purple spots   21 Hoya radicalis
+ Leaf blade obovate to orbicular, 4-8 cm, lateral veins ca. 4 pairs or obscure; flower clusters globose; corolla uniformly white.   (8)
8 (7) Apex of leaf blade retuse or concave, lateral veins obscure   20 Hoya kerrii
+ Apex of leaf blade entire, lateral veins conspicuous, ca. 4 pairs   22 Hoya liangii
9 (6) Leaf blades often pandurate, sometimes oblong; peduncle 0.3 cm; corolla yellow or reddish   27 Hoya pandurata
+ Leaf blade lanceolate or oblong-lanceolate to oblong-oblanceolate; peduncle 1-8 cm; corolla white.   (10)
10 (9) Branchlets and petiole hairy; peduncle 1-4 cm; corolla reflexed; corona lobes erect.   (11)
+ Branchlets and petiole glabrous; peduncle 5-8 cm; corolla rotate; corona lobes spreading-stellate.   (12)
11 (10) Branchlets and petiole setose; lateral veins of leaf blade conspicuous adaxially; inflorescences terminal   25 Hoya lii
+ Branchlets and petiole minutely puberulent at least when young; lateral veins of leaf blade ill defined adaxially; inflorescences extra-axillary   26 Hoya mekongensis
12 (10) Leaf blade 1.5-2 cm wide; flower clusters flat topped; corolla 1-1.5 cm in diam., villous inside   28 Hoya mengtzeensis
+ Leaf blade 2.5-4.5 cm wide; flower clusters globose; corolla ca. 3 cm in diam., obscurely puberulent inside   29 Hoya griffithii
13 (3) Corolla purple; inflorescences on hanging leafless branches (leaves not known)   11 Hoya commutata
+ Corolla white, pink, yellow, or yellowish; inflorescences always on normal leafy branches.   (14)
14 (13) Leaf blade linear, 0.4-0.7 cm wide.   (15)
+ Leaf blade not linear, 1-6.5 cm wide.   (16)
15 (14) Leaf blade 2.5-5 cm, hirsute abaxially; inflorescences terminal; corolla lobes glabrous   31 Hoya linearis
+ Leaf blade 10-20 cm, glabrous; inflorescences extra-axillary; corolla lobes pubescent   30 Hoya longifolia
16 (14) Corolla strongly reflexed; corona distinctly stalked, lobes spurred at base   1 Hoya multiflora
+ Corolla not reflexed; corona sessile, lobes not spurred.   (17)
17 (16) Leaf apex caudate; flowers yellow or yellowish.   (18)
+ Leaf apex obtuse to acuminate; flowers white or reddish.   (20)
18 (17) Leaf blade 4.5-7 cm, base rounded, lateral veins 4 or 5 pairs, obscure; peduncle ca. 0.4 cm; pedicel 1-1.2 cm; corolla lobes glabrous   5 Hoya lasiogynostegia
+ Leaf blade 9-13 cm, base cuneate, lateral veins 10-12 pairs, conspicuous; peduncle 1.5-2.5 cm; pedicel 2-2.5 cm; corolla lobes puberulent inside.   (19)
19 (18) Leaf blade symmetrical at base, lateral veins at right angle to midvein, uniformly colored   3 Hoya fusca
+ Leaf blade oblique at base, lateral veins obliquely ascending, midrib with pale zone on both sides   4 Hoya lipoensis
20 (17) Lateral veins of leaf blade obscure or obsolete.   (21)
+ Lateral veins of leaf blade conspicuous.   (26)
21 (20) Leaf blade oblong or widest above middle.   (22)
+ Leaf blade widest below middle.   (23)
22 (21) Petiole and abaxial surface of leaf blade pubescent; corolla ca. 1.2 cm in diam.   6 Hoya thomsonii
+ Petiole and leaf blade glabrous; corolla 2-4 cm in diam.   30 Hoya longifolia
23 (21) Pedicel longer than peduncle.   (24)
+ Pedicel shorter than or as long as peduncle.   (25)
24 (23) Leaf blade lanceolate to ovate-lanceolate, apex acuminate; peduncle 1-5 cm; sepals puberulent outside   7 Hoya siamica
+ Leaf blade elliptic or elliptic-oblong, apex obtuse; peduncle ca. 0.3 cm; sepals glabrous outside   8 Hoya revolubilis
25 (23) Corolla 0.6-0.7 cm wide, violet dotted, base conspicuously white villous inside, lobes much longer than tube   9 Hoya lacunosa
+ Corolla 1.5-2 cm wide, not violet dotted, densely papillate inside, lobes as long as tube   10 Hoya carnosa
26 (20) Flower clusters flat topped.   (27)
+ Flower clusters globose.   (28)
27 (26) Leaf blade cordate at base; petiole and abaxial side of midvein puberulent; peduncle shorter than pedicel; calyx outside and corolla puberulent   12 Hoya cordata
+ Leaf blade rounded at base; petiole and midvein glabrous; peduncle much longer than pedicel; calyx and corolla glabrous   13 Hoya nervosa
28 (26) Petiole up to 0.4 cm.   (29)
+ Petiole 1-3 cm.   (30)
29 (28) Lateral leaf veins numerous, parallel; peduncle shorter than pedicel; corolla lobes papillate inside   14 Hoya polyneura
+ Lateral leaf veins 4 or 5 pairs, not parallel; peduncle longer than pedicel; corolla mealy inside   15 Hoya dasyantha
30 (28) Leaves pubescent abaxially, especially along veins.   (31)
+ Leaves glabrous.   (32)
31 (30) Leaf blade short acuminate at apex, cuneate at base; calyx glabrous; corolla ca. 1 cm in diam.   16 Hoya silvatica
+ Leaf blade obtuse or rounded at apex, rounded or broadly cuneate at base; calyx puberulent outside; corolla ca. 1.5 cm in diam.   17 Hoya salweenica
32 (30) Lateral leaf veins ca. 7 pairs; peduncle to 5 cm; calyx pubescent outside; corolla densely pale scurfy inside   18 Hoya fungii
+ Lateral leaf veins ca. 4 pairs; peduncle ca. 2 cm; calyx glabrous; corolla sparsely pubescent inside   19 Hoya ovalifolia

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