1 |
Petals and stamens deciduous after anthesis (rarely tardily); shrubs or rarely subshrubs; plant always without dark glands |
|
(2) |
+ |
Petals and stamens persistent after anthesis; herbs; plant very often with dark glands |
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(39) |
|
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|
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2 (1) |
Styles completely united; anthers ± basifixed |
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(3) |
+ |
Styles free or partly united; anthers markedly dorsifixed |
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(6) |
|
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|
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3 (2) |
Sepals 1-3 mm, mostly broadly ovate to oblong, usually obtuse to rounded, without distinct midvein. |
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33 H. geminiflorum |
+ |
Sepals 3.5-10 mm, mostly lanceolate or narrowly elliptic to linear, acute to subacute (or rarely rounded), often with distinct midvein |
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(4) |
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|
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4 (3) |
Stems 4-angled or 4-winged when mature; leaves usually narrowly elliptic to lanceolate, acute to subacute. |
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34 H. subalatum |
+ |
Stems terete when mature; leaves ovate to elliptic or obovate, subacute to rounded |
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(5) |
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5 (4) |
Sepals obovate to lanceolate, 7-10 mm, ascending in fruit; petals 10-20 mm; leaves ovate to elliptic. |
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31 H. formosanum |
+ |
Sepals linear-lanceolate, 3.5-8 mm, reflexed in fruit; petals 20-28 mm; leaves elliptic to obovate. |
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32 H. nakamurae |
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6 (2) |
Leaves (at least upper) sessile, thickly leathery, without visible reticulate venation abaxially; styles free |
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1 H. augustinii |
+ |
Leaves all subsessile or short petiolate or, if sessile, then visibly ± densely reticulate-veined abaxially and/or with styles ± united |
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(7) |
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7 (6) |
Stem prostrate to ascending or pendent, forming low mats or clumps; styles ca. as long as ovary. |
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2 H. reptans |
+ |
Stem erect to straggling, not mat-forming |
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(8) |
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|
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8 (7) |
Styles partly coherent or ± united, at least 1.4 × as long as ovary; leaves usually densely reticulate-veined abaxially (except in |
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8 H. longistylum). |
+ |
Styles free, usually less than 1.4 × as long as ovary; leaves usually laxly or not visibly reticulate-veined abaxially |
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(14) |
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9 |
Styles coherent in lower half at first |
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(10) |
+ |
Styles united nearly to apex; petiole absent or to 1.5 mm (except in |
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4 H. monogynum). |
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|
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10 (9) |
Leaves sessile, blade ovate-oblong to ovate, venation densely reticulate. |
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3 H. griffithii |
+ |
Leaf petiole 1.5-2 mm, blade oblong to elliptic, venation laxly reticulate. |
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5 H. cohaerens |
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11 |
Suffrutex; petals and stamens tardily deciduous. |
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7 H. elatoides |
+ |
Shrub; petals and stamens soon deciduous |
|
(12) |
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|
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12 (11) |
Leaves without visible reticulate venation abaxially, 1-3.3 cm; inflorescence always 1-flowered. |
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8 H. longistylum |
+ |
Leaves densely reticulate-veined abaxially, (2-)3-11 cm; inflorescence 1-30-flowered, terminal on long shoots |
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(13) |
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13 (12) |
Leaves with base cuneate to rounded, or if cordate then apex rounded; leaf blade usually broadest at or above middle. |
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4 H. monogynum |
+ |
Leaves with base cordate-amplexicaul, apex acute to acuminate; leaf blade elliptic-ovate to broadly ovate. |
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6 H. prattii |
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14 (8) |
Leaves sessile, densely reticulate-veined. |
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3 H. griffithii |
+ |
Leaves subsessile or petiolate, laxly or not visibly reticulate-veined |
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(15) |
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|
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15 (14) |
Petals waxy, golden yellow; stamens 0.25-0.35 × as long as petals, densely crowded |
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(16) |
+ |
Petals not waxy, golden to pale yellow; stamens longer than 0.35 × as long as petals or, if equal or shorter, then not densely crowded |
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(18) |
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16 (15) |
Sepals spreading-incurved in bud and in fruit, often becoming markedly ribbed; capsule ovoid to ovoid-conic. |
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11 H. hookerianum |
+ |
Sepals erect to spreading in bud, spreading to recurved in fruit, not ribbed; capsule narrowly ovoid-cylindric to narrowly ellipsoid-cylindric |
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(17) |
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17 (16) |
Leaves narrowly ovate to lanceolate, base rounded, with dense abaxial glands and short laminar glandular streaks; sepals spreading in bud, reflexed in fruit. |
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9 H. fosteri |
+ |
Leaves narrowly lanceolate, base cuneate, without abaxial glands, with long laminar glandular streaks; sepals erect in bud, spreading in fruit. |
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10 H. wardianum |
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|
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18 (15) |
Sepals spreading to recurved at anthesis and in fruit; leaf venation often markedly reticulate or at least conspicuous abaxially (except in |
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13 H. acmosepalum). |
+ |
Sepals erect and outcurving or appressed in flower and fruit; leaf venation neither visibly reticulate nor conspicuous |
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(25) |
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19 |
Leaves elliptic to oblong or (lower) oblanceolate (broadest at or above middle), never triangular; sepals deciduous or outcurving |
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(20) |
+ |
Leaves ovate or lanceolate to oblong-lanceolate (broadest below middle), sometimes triangular; sepals persistent, spreading to recurved |
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(21) |
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20 (19) |
Leaves subsessile, narrowly elliptic, markedly reticulate-veined; sepals deciduous, spreading, ovate to elliptic, often leaflike; capsule 16-21 mm, purplish red when maturing. |
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12 H. subsessile |
+ |
Leaves petiolate (0.5-1.5 mm), narrowly elliptic to oblong, not reticulate-veined but with conspicuous intramarginal vein; sepals persistent, ovate to narrowly lanceolate, not leaflike; capsule 9-15 mm, bright red at first when maturing. |
|
13 H. acmosepalum |
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21 (19) |
Sepals elliptic to oblong or foliose; leaves lanceolate to ovate-lanceolate, ± triangular, with venation (secondary and sometimes tertiary) conspicuous |
|
(22) |
+ |
Sepals lanceolate-elliptic to lanceolate (or rarely leaflike); leaves lanceolate to oblong or ovate, not triangular, usually without conspicuous venation |
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(23) |
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|
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22 (21) |
Petals obovate-oblanceolate, flowers stellate; stamens ca. 3/5 as long as petals; styles 0.85-1 × as long as ovary; leaves densely glaucous abaxially, petiole 0.5-2 mm. |
|
14 H. maclarenii |
+ |
Petals broadly obovate to obovate-suborbicular, flowers shallowly to deeply cupped; stamens 0.35-0.4 × as long as petals; styles 0.35-0.7 × as long as ovary; leaves thinly or not glaucous abaxially, petiole 2-4 mm. |
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15 H. choisyanum |
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23 (21) |
Styles shorter than ovary; inflorescence 1(-3)-flowered; leaves triangular-lanceolate to triangular-ovate. |
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18 H. curvisepalum |
+ |
Styles ca. as long as ovary or more; inflorescence (1-)3-14-flowered; leaves oblong-lanceolate or lanceolate to narrowly ovate |
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(24) |
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24 (23) |
Styles outcurved distally; inflorescence branches usually relatively stout; leaf abaxial glands usually few or none. |
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16 H. lancasteri |
+ |
Styles straight or flexuous; inflorescence branches relatively slender; leaf abaxial glands dense. |
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17 H. stellatum |
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25 (18) |
Sepals with broad hyaline margin; stems erect to arching with ± numerous lateral branches, often frondose, markedly ancipitous (2-edged) when young |
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(26) |
+ |
Sepals without or with a very narrow hyaline margin; stems erect to spreading but not frondose, not or scarcely ancipitous (2-edged) when young |
|
(30) |
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|
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26 (25) |
Sepals entire, at least outer broadly elliptic or broadly oblong to circular, obtuse to rounded; stems erect, not frondose. |
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19 H. henryi |
+ |
Sepals eroded-denticulate or, if entire, then elliptic or narrowly oblong to oblanceolate, rounded to acute; stems erect or spreading, sometimes frondose |
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(27) |
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|
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27 (26) |
Sepals eroded-denticulate to subentire, usually apiculate, broadly elliptic to broadly ovate; leaf apex usually apiculate-obtuse to rounded |
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(28) |
+ |
Sepals entire, rarely apiculate, elliptic to narrowly oblong or obovate-spatulate; leaf apex acute to rounded-apiculate |
|
(29) |
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|
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28 (27) |
Stems erect to arching or rarely divaricate, not frondose, ± persistently 4-lined; leaf-apex acute to rounded but rarely apiculate; capsule 10-14 mm. |
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19 H. henryi |
+ |
Stems spreading, sometimes frondose, soon 2-lined; leaf-apex obtuse to rounded, always apiculate; capsule 9-11 mm. |
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20 H. patulum |
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|
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29 (27) |
Flower buds acute to obtuse; sepals subacute to rounded; stems erect or arching, not or only distally frondose. |
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19 H. henryi |
+ |
Flower buds obtuse to rounded; sepals rounded; stems arching, often frondose. |
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21 H. uralum |
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|
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30 (25) |
Stems slender, spreading especially when young; capsules lobed or grooved |
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(31) |
+ |
Stems stout, erect to arching but not spreading; capsules not lobed or grooved |
|
(32) |
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|
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31 (30) |
Styles 0.5-0.85 × as long as ovary; sepals with inconspicuous midvein, outcurving in fruit; flower buds ovoid to ovoid-pyramidal, obtuse to acute. |
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22 H. lagarocladum |
+ |
Styles 1.5-1.8 × as long as ovary; sepals with conspicuous midvein, erect in fruit; flower buds narrowly ovoid, acute to acuminate. |
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23 H. wilsonii |
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32 (30) |
Styles 1.2-1.35 × as long as ovary; petals spreading to reflexed, apiculus acute; sepals acute, spreading in fruit. |
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28 H. kouytchense |
+ |
Styles 0.6-1.1 × as long as ovary; petals incurved to spreading, apiculus obtuse to rounded; sepals acute to rounded, spreading to erect in fruit |
|
(33) |
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33 (32) |
Sepals ± spreading in bud and sometimes in fruit; ovary and capsule narrowly cylindric-ellipsoid to broadly pyramidal-ovoid; leaf apex acute to rounded-apiculate. |
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24 H. beanii |
+ |
Sepals erect in bud and erect or incurved in fruit; ovary and capsule ovoid to subglobose or pyramidal-ovoid; leaf apex obtuse to rounded |
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(34) |
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|
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34 (33) |
Leaves broadly elliptic or very broadly elliptic-oblong to orbicular, margin plane |
|
24 H. beanii |
+ |
Leaves lanceolate to broadly ovate or, if circular, then margin undulate |
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(35) |
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35 (34) |
Branches spreading; sepals acute to rounded-apiculate; stamens ca. 0.4 × as long as petals; leaves with lower 1 or 2 pairs of main lateral veins usually free. |
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25 H. addingtonii |
+ |
Branches erect to arching; sepals acute to rounded; stamens ca. 0.5 × as long as petals or more, or, if relatively shorter, then sepals rounded; leaves with all main lateral veins incurved to join next upper vein |
|
(36) |
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36 (35) |
Ovary and capsule subglobose to broadly or narrowly ovoid; leaves lanceolate to ovate, rhombic, or subcircular; sepal margin not or scarcely hyaline |
|
(37) |
+ |
Ovary and capsule pyramidal-ovoid; leaves triangular-lanceolate or lanceolate-oblong to ± broadly triangular-ovate; sepal margin distinctly hyaline |
|
(38) |
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|
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37 (36) |
Sepals ovate to broadly elliptic, obtuse or apiculate to rounded; capsule broadly ovoid; flowers 4-6 cm in diam.; stamens (0.5-)0.6-0.7 × as long as petals; leaf ratio (l:w) = 1.8-2.5, base cuneate to rounded, margin plane. |
|
26 H. latisepalum |
+ |
Sepals narrowly elliptic or oblong to obovate, usually rounded; capsule narrowly ovoid; flowers 2.5-3.5 cm in diam.; stamens 0.35-0.45(-0.6) × as long as petals; leaf ratio (l:w) = 1.1-1.6, base truncate to subcordate or, if cuneate, then margin undulate. |
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27 H. bellum |
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38 (36) |
Sepals acute to obtuse, outcurved in fruit; stems ± persistently 4-lined; leaves usually lanceolate or lanceolate-oblong to ovate-oblong; petals spreading to reflexed; flower buds subacute to apiculate. |
|
29 H. pseudohenryi |
+ |
Sepals rounded or rarely apiculate, erect in fruit; stems soon terete; leaves triangular-ovate or, more rarely, lanceolate or ± broadly ovate; petals incurved; flower buds rounded. |
|
30 H. forrestii |
|
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|
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39 (1) |
Styles (4 or)5; stamen fascicles 5, free; dark glands absent |
|
(40) |
+ |
Styles (2 or)3(or 4); stamen fascicles usually apparently 3 or stamens irregularly arranged; dark glands usually present |
|
(41) |
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|
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40 (39) |
Leaf apex acute to obtuse, blade mostly lanceolate to elliptic-oblong or linear; mature stem internodes sharply 4-lined to narrowly 4-winged. |
|
35 H. ascyron |
+ |
Leaf apex rounded to shallowly retuse, blade oblong to oblong-lanceolate, oblong-ovate, or triangular-oblong; mature stem internodes partially 2- or 4-lined or terete. |
|
36 H. przewalskii |
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|
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41 (39) |
Dark glands absent; stem internodes 4-lined or narrowly 4-winged; stamens irregularly arranged |
|
(42) |
+ |
Dark (black or reddish) glands present on at least sepals and/or anthers or leaves; stem internodes 2-lined or terete; stamens usually apparently 3(or 4)-fascicled |
|
(43) |
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|
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42 (41) |
Stamens 30-40, styles 0.7-1.8 mm; stems branching strictly from base or unbranched, not rooting; leaves ovate-lanceolate to linear (l:w = 3-5). |
|
63 H. gramineum |
+ |
Stamens 5-30, styles 0.4-1 mm; stems usually branching ± widely along most of stem, rooting at base; leaves usually ovate or triangular-ovate to oblong or elliptic (l:w = 1.5-2.5). |
|
64 H. japonicum |
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|
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43 (41) |
Anther gland amber; dark glands sometimes present on leaves but not forming inframarginal row; seeds with testa ± markedly papillose |
|
(44) |
+ |
Anther gland dark; dark glands nearly always forming inframarginal row on leaves; seeds with testa reticulate to foveolate |
|
(46) |
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|
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44 (43) |
Stem and leaves pubescent; inflorescence usually pyramidal; sepals gland-fringed; stems creeping and rooting. |
|
62 H. hirsutum |
+ |
Stem and leaves glabrous; inflorescence ± cylindric or flat-topped; sepal margin with few or no glands; stems not creeping or rooting |
|
(45) |
|
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|
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45 (44) |
Inflorescence ± cylindric; stem smooth. |
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60 H. elongatum |
+ |
Inflorescence flat-topped; stem scabrid with red-gland-tipped emergences. |
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61 H. scabrum |
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46 (43) |
Stem internodes markedly 2(or 4)-lined |
|
(47) |
+ |
Stem internodes terete or rarely 2-4(-6)-lined when stems weak or slender |
|
(50) |
|
|
|
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47 (46) |
Capsule valves with central linear and lateral vesicular glands; stems much branched distally with branching divergent-ascending and black glands confined to lines. |
|
57 H. perforatum |
+ |
Capsule valves with linear glands only; stems sparingly branched below inflorescence with branching virgate and black glands not confined to lines or completely absent |
|
(48) |
|
|
|
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48 (47) |
Stems with dark glands, erect; capsule broadly ovoid to narrowly conic; stamens apparently 3-fascicled. |
|
56 H. attenuatum |
+ |
Stems without dark glands, suberect to ascending; capsule broadly to narrowly ovoid; stamens irregularly arranged |
|
(49) |
|
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|
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49 (48) |
Sepals without dark laminar streaks or, if present, then apex obtuse; leaves variable in size and shape but, if less than 15 mm, then narrow (l:w = 4-12). |
|
58 H. nagasawae |
+ |
Sepals with dark laminar streaks, apex acute to subacuminate; leaves short (4-12 mm) and broad (l:w = 2-3.1). |
|
59 H. nokoense |
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|
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50 (46) |
Leaf pairs perfoliate; capsule valves with vesicular glands only. |
|
37 H. sampsonii |
+ |
Leaf pairs free; capsule valves with longitudinal vittae only |
|
(51) |
|
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|
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51 (50) |
Leaves and sometimes sepals and petals with laminar glands dark only |
|
(52) |
+ |
Leaves, sepals, and petals with laminar glands (if present) partly or completely pale |
|
(55) |
|
|
|
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52 (51) |
Sepals entire, sepals and petals 5 |
|
(53) |
+ |
Sepals glandular-ciliate or, if entire, then sepals and petals 4 |
|
(54) |
|
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|
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53 (52) |
Leaves sessile, triangular-ovate to narrowly elliptic, not minutely papillose; stems usually unbranched below inflorescence. |
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55 H. erectum |
+ |
Leaves petiolate, oblong to obovate, adaxially minutely papillose; stems branching from most nodes. |
|
53 H. enshiense |
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|
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54 (52) |
Styles 4-5 mm; capsule broadly ovoid; uppermost leaf pair bracteose; sepals and petals 5. |
|
47 H. trigonum |
+ |
Styles 1.5-3 mm; capsule globose; uppermost leaf foliar; sepals and petals 4. |
|
50 H. monanthemum |
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55 (51) |
Sepals entire, oblong to ovate-oblong, obtuse (or rarely acute) to rounded, not leaflike |
|
(56) |
+ |
Sepals dark-gland-fringed or, if entire, then acute or leaflike |
|
(58) |
|
|
|
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56 (55) |
Leaf laminar glands mostly black. |
|
53 H. enshiense |
+ |
Leaf laminar glands all or mostly pale or absent |
|
(57) |
|
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|
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57 (56) |
Leaves without or rarely with a few large pale or pale and dark laminar glands, main lateral veins 2- or 3-paired. |
|
52 H. faberi |
+ |
Leaves with dense small pale laminar glands; main lateral veins 3-4-paired. |
|
54 H. taihezanense |
|
|
|
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58 (55) |
Leaves with laminar glands large, prominent, dense; leaf blade usually lanceolate or narrowly elliptic to linear or obovate; marginal glands round-topped |
|
(59) |
+ |
Leaves with laminar glands small, not prominent, often sparse, or absent; leaf blade ovate or triangular-ovate to elliptic, oblong, or circular; marginal glands flat-topped |
|
(65) |
|
|
|
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59 (58) |
Sepals entire or occasionally with few glandular cilia; bracts without or with short glandular auricles; leaves entire |
|
(60) |
+ |
Sepals regularly glandular-ciliate; bracts and sometimes upper leaves glandular-ciliate and with glandular-ciliate auricles |
|
(62) |
|
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60 (59) |
Leaves with petiole 1-10 mm, base usually cuneate to angustate; sepals with few irregularly spaced marginal glands or none; styles 1-2.5 mm, 0.7-1.3 × as long as ovary. |
|
39 H. petiolulatum |
+ |
Leaves sessile or with broad petiole to 1 mm, base subcordate-amplexicaul to cuneate; sepals with marginal glands in ± regular row; styles 2.5-10 mm, 1.5-3 × as long as ovary |
|
(61) |
|
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|
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61 (60) |
Leaves oblong-lanceolate to narrowly oblong, 5-13 mm wide; sepal and petal laminar glands pale. |
|
38 H. seniawinii |
+ |
Leaves oblong-ovate, 10-16 mm wide; sepal and petal laminar glands black. |
|
40 H. hubeiense |
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|
|
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62 (59) |
Leaves broadly ovate-oblong or broadly elliptic to suborbicular, 6-14 mm, entire, apex rounded, without glandular auricles. |
|
44 H. kingdonii |
+ |
Leaves lanceolate to oblong-lanceolate or linear, 1-6 cm, sometimes glandular-ciliate, apex acute to obtuse, the upper with glandular auricles |
|
(63) |
|
|
|
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63 (62) |
Styles ca. 1.7 mm, shorter than ovary; sepals ovate, 3-5 mm; leaves ternate. |
|
43 H. austroyunnanicum |
+ |
Styles 3-18 mm, longer than ovary; sepals lanceolate to oblong, 5-9 mm; leaves paired |
|
(64) |
|
|
|
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64 (63) |
Styles 3-7 × as long as ovary, 6-18 mm; pedicels 0.5-3(-6) mm; petal laminar glands pale. |
|
41 H. hengshanense |
+ |
Styles ca. 2 × as long as ovary, 3-8 mm; pedicels 3-12 mm; petal laminar glands all or mostly dark. |
|
42 H. elodeoides |
|
|
|
|
65 (58) |
Leaves elliptic-oblong to lanceolate and/or with glandular-ciliate margin and auricles |
|
(66) |
+ |
Leaves (triangular-)ovate to elliptic or oblong or obovate to oblanceolate, entire (uppermost pair sometimes bracteose) |
|
(67) |
|
|
|
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66 (65) |
Ovary and capsule ellipsoid to cylindric-ellipsoid; leaves all sessile, entire; sepals lanceolate to narrowly oblong-elliptic, acute, glandular-ciliate. |
|
45 H. daliense |
+ |
Ovary and capsule ovoid to globose; leaves (at least upper) glandular-ciliate, lower often petiolate; sepals broadly oblong or ± broadly elliptic, acute to obtuse, glandular-ciliate to -laciniate. |
|
46 H. wightianum |
|
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|
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67 (65) |
Ovary and capsule ovoid (or ovary ellipsoid) to subglobose; styles 1.5-3 mm; uppermost leaf pair leaflike |
|
(68) |
+ |
Ovary and capsule ± ellipsoid or, if ovoid-ellipsoid to broadly ovoid, then styles 4-5 mm; uppermost leaf pair bracteose |
|
(69) |
|
|
|
|
68 (67) |
Leaves (at least uppermost) sessile; sepals with pale laminar glandular lines or, if dark, then margin glandular-ciliate; ovary ovoid to subglobose. |
|
50 H. monanthemum |
+ |
Leaves all short petiolate; sepals all with dark linear laminar glands, entire; ovary ellipsoid. |
|
51 H. subcordatum |
|
|
|
|
69 (67) |
Styles 4-5 mm; capsule broadly ovoid; leaves all sessile, blade oblong to triangular-ovate. |
|
47 H. trigonum |
+ |
Styles 2-3 mm; capsule ± ellipsoid; some or all leaves petiolate, blade triangular-ovate to elliptic or oblanceolate to obovate |
|
(70) |
|
|
|
|
70 (69) |
Stem unbranched below apical node or with a few short suberect branches; styles usually longer than ovary; sepals usually without dark laminar glands. |
|
48 H. ludlowii |
+ |
Stem nearly always with long ± spreading branches from up to 5 nodes; styles usually (?always) ca. as long as ovary or less; sepals usually with dark laminar glands. |
|
49 H. himalaicum |