1 |
Fruiting calyx with 5, equal or subequal straight teeth. |
|
(2) |
+ |
Fruiting calyx with subequal teeth or 2-lipped, recurved. |
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(13) |
|
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|
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2 (1) |
Fruiting calyx tubular or tubular-campanulate to ovoid; inflorescences pyramidal paniculate. |
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(3) |
+ |
Fruiting calyx campanulate to broadly campanulate; inflorescences loosely paniculate. |
|
(8) |
|
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3 (2) |
At least some leaves in whorls of 3 or 4 |
|
1 Isodon ternifolius |
+ |
Leaves always opposite. |
|
(4) |
|
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4 (3) |
Stems and inflorescences densely, spreading, purplish or brownish, villous-velutinous or softly tomentose; leaves corrugate. |
|
(5) |
+ |
Stems and inflorescences with indumentum not as above; leaves not or only slightly rugulose. |
|
(6) |
|
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|
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5 (4) |
Leaves ovate to triangular-ovate, 1.5-7 × 1-4 cm; petiole 2-8 mm; corolla purple to whitish blue |
|
2 Isodon enanderianus |
+ |
Leaves triangular-ovate, 1-2.5 × 1-2.5 cm; petiole 15-25 mm; corolla white |
|
3 Isodon brevifolius |
|
|
|
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6 (4) |
Leaves long petiolate, slightly pubescent along midribs and veins; panicles without floral leaves; calyx gray lanate, glabrescent and tubular in fruit |
|
4 Isodon eriocalyx |
+ |
Leaves sessile or short petiolate; inflorescence and calyx not as above. |
|
(7) |
|
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|
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7 (6) |
Leaves ovate, sometimes cordate, 1.5-5 × 0.8-4.5 cm; adaxially puberulent or finely pilose; abaxially gray tomentose; fruiting calyx densely pilose, ovoid |
|
5 Isodon phyllostachys |
+ |
Leaves broadly ovate, 7-17 cm, ca. 1/2 as wide as long; abaxially densely strigose, sparsely glandular; calyx pubescent |
|
6 Isodon pantadenius |
|
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8 (2) |
Leaves lanceolate to narrowly lanceolate; calyx teeth lanceolate; nutlets apex pubescent |
|
7 Isodon nervosus |
+ |
Leaves broadly ovate to ovate-lanceolate; calyx teeth triangular; nutlets glabrous. |
|
(9) |
|
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|
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9 (8) |
Fruiting calyx as long as wide; nutlets glandular |
|
8 Isodon amethystoides |
+ |
Fruiting calyx longer than wide; nutlets not glandular. |
|
(10) |
|
|
|
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10 (9) |
Terminal panicle few flowered, ca. 2 cm |
|
9 Isodon websteri |
+ |
Terminal panicle spreading, many flowered. |
|
(11) |
|
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|
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11 (10) |
Adaxial leaf surface sparsely puberulent, abaxial surface whitish pilose along midrib and veins; calyx with minute hairs, glabrescent especially in fruit |
|
10 Isodon inflexus |
+ |
Leaves sparsely puberulent, glandular or glabrescent; calyx densely pubescent. |
|
(12) |
|
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|
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12 (11) |
Leaves ovate to ovate-lanceolate, ± glabrescent, apex subacuminate, base cuneate, coarsely incurved, serrate; calyx densely whitish pubescent, teeth equal to calyx tube in length; stamens and style included; nutlets apex whitish barbate |
|
11 Isodon serra |
+ |
Leaves ovate to broadly ovate, sparsely puberulent, glandular; apical tooth ovate to lanceolate, apex acuminate, base broadly cuneate to abruptly attenuate, acutely serrate to crenate-dentate; calyx densely appressed whitish pubescent, bluish (especially in var. glaucocalyx), teeth shorter than calyx tube; stamens and style exserted; nutlets apex verruculose |
|
12 Isodon japonicus |
|
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|
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13 (1) |
Cymes few flowered, axillary, in widely spaced verticillasters or in terminal racemose panicles; corolla tube conspicuously pouched or spurred on upper side near base; fruiting calyx conspicuously 2-lipped. |
|
(14) |
+ |
Cymes loosely many flowered, often conspicuously pedunculate, rarely in a dense verticillaster; corolla tube subequal or saccate, never short spurred on upper side near base; fruiting calyx with 5 subequal teeth or 2-lipped. |
|
(18) |
|
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14 (13) |
Leaves narrowly lanceolate to oblong, 3 × or more longer than broad. |
|
(15) |
+ |
Leaves much narrower, 2 × or less longer than broad. |
|
(16) |
|
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15 (14) |
Corolla to 8 mm, tube conspicuously pouched, to 2 mm in diam. on upper side near base, abruptly declinate above base, slightly constricted, ca. 1.2 mm in diam. at throat |
|
73 Isodon gibbosus |
+ |
Corolla ca. 7 mm, tube pouched on upper side near base, not abruptly declinate above base, throat subequal to tube |
|
74 Isodon lungshengensis |
|
|
|
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16 (14) |
Cymes in axils of mid and upper cauline leaves not reduced upwards, in many widely spaced verticillasters |
|
75 Isodon melissoides |
+ |
Cymes in axils of floral leaves gradually reduced toward branch apex, in terminal spicate panicles. |
|
(17) |
|
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|
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17 (16) |
Petiole to 4.5 cm; corolla tube abruptly spurred on upper part near base, spur ca. 3 mm wide, conspicuously exserted, throat ca. 1.5 mm wide |
|
76 Isodon brevicalcaratus |
+ |
Petiole 0.2-1 cm; corolla tube saccate on upper part above base, tube equal in width elsewhere, ca. 2 mm in diam. at throat |
|
77 Isodon adenanthus |
|
|
|
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18 (13) |
Corolla 1.2-2.0 cm, tube 2 × or more longer than limb. |
|
(19) |
+ |
Corolla less than 1.2 cm, to 2 × as long as limb if tube stout. |
|
(24) |
|
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|
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19 (18) |
Fruiting calyx conspicuously 2-lipped; corolla tube ca. 3 × longer than limb |
|
13 Isodon longitubus |
+ |
Fruiting calyx inconspicuously 2-lipped; corolla tube less than 3 × longer than limb. |
|
(20) |
|
|
|
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20 (19) |
Plants less than 30 cm tall; cymes 1-3-flowered, in narrow terminal panicles |
|
14 Isodon gesneroides |
+ |
Plants more than 30 cm tall; cymes 3- to many flowered, in ± spreading panicles. |
|
(21) |
|
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|
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21 (20) |
Calyx broadly campanulate, purplish or dark purple-red. |
|
(22) |
+ |
Calyx campanulate, not tinged purplish. |
|
(23) |
|
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22 (21) |
Panicles strongly spreading, to 30 × 8-9 cm; pedicels 2-7 mm |
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15 Isodon forrestii |
+ |
Panicles slightly spreading, (6-)10-15(-20) cm; floral leaves gradually becoming small bracts at apex; pedicels 1-3 mm |
|
16 Isodon irroratus |
|
|
|
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23 (21) |
Leaves rhombic, coarsely crenate-serrate above middle; petioles to 3.5 cm |
|
17 Isodon wardii |
+ |
Leaves narrowly ovate, crenate-dentate above basal 1/3; petioles less than 1 cm |
|
18 Isodon bulleyanus |
|
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|
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24 (18) |
Fruiting calyx conspicuously 2-lipped. |
|
(25) |
+ |
Fruiting calyx with 5 equal or subequal teeth or inconspicuously 2-lipped. |
|
(41) |
|
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25 (24) |
Calyx lips extending to middle of calyx. |
|
(26) |
+ |
Calyx lips extending below middle of calyx. |
|
(31) |
|
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26 (25) |
Cymes in narrow panicles. |
|
(27) |
+ |
Cymes in spreading panicles. |
|
(29) |
|
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27 (26) |
Leaf blade base cordate; panicles secund |
|
58 Isodon secundiflorus |
+ |
Leaf blade base not cordate; panicles not secund. |
|
(28) |
|
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28 (27) |
Stamens included; lower corolla lip usually longer than corolla tube |
|
56 Isodon coetsa |
+ |
Stamens slightly exserted; lower corolla lip shorter than corolla tube |
|
57 Isodon phyllopodus |
|
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29 (26) |
Leaf blade base cordate |
|
59 Isodon sculponeatus |
+ |
Leaf blade base broadly cuneate to subcuneate, never cordate. |
|
(30) |
|
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|
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30 (29) |
Terminal panicles to 30 × 20 cm, of 5-11-flowered cymes |
|
60 Isodon megathyrsus |
+ |
Terminal panicles 6-10(-15) × ca. 2.5 cm, of 3-5-flowered cymes |
|
61 Isodon henryi |
|
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|
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31 (25) |
Cymes in spreading panicles. |
|
(32) |
+ |
Cymes in racemose or falsely racemose narrow panicles. |
|
(35) |
|
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|
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32 (31) |
Leaf blade base ± shallow cordate. |
|
(33) |
+ |
Leaf blade base not cordate. |
|
(34) |
|
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33 (32) |
Teeth of fruiting calyx broadly triangular, apex acute; filaments basally pilose |
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69 Isodon scrophularioides |
+ |
Teeth of fruiting calyx triangular to lanceolate-triangular, apex acuminate; filaments bearded below middle |
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70 Isodon flabelliformis |
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34 (32) |
Stem shallowly grooved, densely retrorse puberulent; leaves very broadly ovate to oblate-ovate, apex rounded, margin crenate-serrate, apical tooth acute; corolla reddish purple or white |
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71 Isodon latifolius |
+ |
Stem strongly grooved, puberulent on angles otherwise glabrous with scattered glands; leaves broadly ovate to ovate-orbicular, apex acuminate, margin coarse serrate or double serrate, apical tooth acuminate; corolla white |
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72 Isodon weisiensis |
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35 (31) |
Leaves crenate-serrate. |
|
(36) |
+ |
Leaves coarsely serrate, dentate, or serrate-dentate. |
|
(38) |
|
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36 (35) |
Corolla less than 5.5 mm |
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66 Isodon rosthornii |
+ |
Corolla more than 5.5 mm. |
|
(37) |
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37 (36) |
Stem densely retrorse white puberulent along angles; corolla gray or reddish, to 10 mm |
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67 Isodon liangshanicus |
+ |
Stem appressed pubescent; corolla purple or purple-red, ca. 8 mm |
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68 Isodon macrocalyx |
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38 (35) |
Inflorescences racemose or falsely racemose (in latter case basal cymes of inflorescence each with 3 flowers and a short peduncle), terminal and axillary, elongated, 8-20 cm |
|
62 Isodon racemosus |
+ |
Inflorescences ± conspicuously narrow paniculate. |
|
(39) |
|
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39 (38) |
Leaf apex long acuminate, sometimes ± excised |
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63 Isodon excisoides |
+ |
Leaf apex not long acuminate, excised. |
|
(40) |
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40 (39) |
Plants densely white pilose; leaves dentate-serrate |
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64 Isodon albopilosus |
+ |
Plants sparsely white puberulent; leaves coarsely serrate |
|
65 Isodon grosseserratus |
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41 (24) |
Leaf apex conspicuously excised. |
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(42) |
+ |
Leaf apex not excised. |
|
(43) |
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42 (41) |
Abaxial leaf surface densely hairy; apical leaf tooth narrowly lanceolate, 2-2.5 cm, slightly cuneate, not extending to midrib of blade at base; cymes 10-15-flowered |
|
19 Isodon macrophyllus |
+ |
Abaxial leaf surface sparsely hairy or subglabrescent; apical leaf tooth caudate, 4-6 cm, cuneate, extending to midrib of blade at base, entire or 1- to few serrate; cymes (1-)3-5-flowered |
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20 Isodon excisus |
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43 (41) |
Fruiting calyx with 5 short teeth to 1/3 as long as fruiting calyx tube; corolla tube spreading or straight. |
|
(44) |
+ |
Fruiting calyx with 5 long teeth more than 1/3 as long as fruiting calyx tube; corolla tube declinate. |
|
(49) |
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44 (43) |
Corolla mostly yellow or yellowish, rarely rose |
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21 Isodon flavidus |
+ |
Corolla purplish, blue, rose, or white. |
|
(45) |
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45 (44) |
Inflorescences gray tomentulose |
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22 Isodon calcicolus |
+ |
Inflorescences with hairs not as above, hairs never gray. |
|
(46) |
|
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46 (45) |
Leaves narrowly lanceolate to elliptic-lanceolate, base narrowly cuneate, margin ± remotely serrate above middle |
|
23 Isodon walkeri |
+ |
Leaves ovate to ovate-lanceolate, base rounded to cuneate-attenuate, margin crenate or serrate except at base. |
|
(47) |
|
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47 (46) |
Inflorescences papillate velutinuous; leaves scaly hirsute especially on midrib and veins of abaxial surface |
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24 Isodon yuennanensis |
+ |
Inflorescences not papillate velutinous; leaves villous or hirtellous especially on midrib and veins of abaxial surface. |
|
(48) |
|
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48 (47) |
Calyx densely villous |
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25 Isodon hispidus |
+ |
Calyx sparsely villous basally |
|
26 Isodon lophanthoides |
|
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49 (43) |
Calyx teeth extending to middle of calyx or more. |
|
(50) |
+ |
Calyx with small teeth never extending to middle of calyx. |
|
(55) |
|
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50 (49) |
Cymes in interrupted spikelike panicles |
|
50 Isodon interruptus |
+ |
Cymes in ± continuous narrow spikes. |
|
(51) |
|
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51 (50) |
Calyx teeth apex filiform-acuminate |
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51 Isodon barbeyanus |
+ |
Calyx teeth apex never filiform-acuminate. |
|
(52) |
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52 (51) |
Leaves subcircular to ovate-oblong, 1.2-1.8 × 0.6-1.8 cm, strongly netted |
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52 Isodon scoparius |
+ |
Leaves larger, inconspicuously netted. |
|
(53) |
|
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53 (52) |
Floral leaves bractlike; calyx teeth ovate-triangular |
|
53 Isodon muliensis |
+ |
Floral leaves ± similar to stem leaves; calyx teeth narrowly lanceolate to lanceolate-triangular. |
|
(54) |
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54 (53) |
Panicles terminal, 1.5-13 cm, cymes 3-5(-11)-flowered, peduncle 2-5(-9) mm |
|
54 Isodon setschwanensis |
+ |
Panicles terminal or axillary, to 20(-34) cm, cymes 3-15(-23)-flowered, peduncle 4-25(-48) mm |
|
55 Isodon angustifolius |
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55 (49) |
Branchlets, leaves, and calyces sparsely hairy; leaves usually not corrugate. |
|
(56) |
+ |
Branchlets, leaves, and calyces variously dense hairy; leaves mostly corrugate. |
|
(62) |
|
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56 (55) |
Stems ascending; branches and petioles densely pilose; stem leaves triangular-ovate, petiole 1.5-3 cm |
|
43 Isodon flexicaulis |
+ |
Stems erect; branches and petioles sparsely pilose; stem leaves broadly ovate to oblong-lanceolate, petiole less than 1.5 cm. |
|
(57) |
|
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57 (56) |
Stem leaves less than 2 cm. |
|
(58) |
+ |
Stem leaves more than 2 cm. |
|
(60) |
|
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58 (57) |
Stem leaves broadly truncate-ovate to subcircular, glandular puberulent |
|
44 Isodon glutinosus |
+ |
Stem leaves mostly ovate, subrhombic-ovate, to oblong-lanceolate, sometimes broadly ovate, not glandular hairy. |
|
(59) |
|
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59 (58) |
Corolla white, ca. 6 mm, tube included; calyx densely white pilose inside on teeth |
|
45 Isodon medilungensis |
+ |
Corolla purplish, ca. 9 mm, tube exserted; calyx glabrous inside |
|
46 Isodon pharicus |
|
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60 (57) |
Stems subcylindric, densely glandular pubescent; calyx teeth triangular, apex conspicuously mucronate |
|
47 Isodon mucronatus |
+ |
Stems obtusely 4-angled, pubescent or glandular pubescent; calyx teeth ovate-triangular, apex acute, never mucronate. |
|
(61) |
|
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61 (60) |
Branchlets and petioles densely gray pubescent, not glandular pubescent |
|
48 Isodon silvaticus |
+ |
Branchlets and petioles densely glandular pubescent |
|
49 Isodon pleiophyllus |
|
|
|
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62 (55) |
Branchlets, leaves, and calyces densely stellate tomentose or stellate lanate. |
|
(63) |
+ |
Branchlets, leaves, and calyces with different hairs than above. |
|
(68) |
|
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|
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63 (62) |
Calyx teeth ca. 1/3 as long as calyx tube. |
|
(64) |
+ |
Calyx teeth ca. as long as calyx tube. |
|
(66) |
|
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|
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64 (63) |
Leaves without a brown gland on lower surface of each tooth apex |
|
29 Isodon rugosus |
+ |
Leaves with a brown gland on lower surface of each tooth apex. |
|
(65) |
|
|
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65 (64) |
Panicles 8-20 cm |
|
27 Isodon leucophyllus |
+ |
Panicles 3-5 cm |
|
28 Isodon adenolomus |
|
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66 (63) |
Leaves 3-8 cm |
|
30 Isodon grandifolius |
+ |
Leaves less than 2.5 cm. |
|
(67) |
|
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|
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67 (66) |
Leaf blade base rounded to shallowly cordate |
|
31 Isodon oresbius |
+ |
Leaf blade base cuneate to truncate-cuneate |
|
32 Isodon dawoensis |
|
|
|
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68 (62) |
Leaves conspicuously discolored. |
|
(69) |
+ |
Leaves not discolored. |
|
(73) |
|
|
|
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69 (68) |
Calyx teeth narrowly triangular or triangular-lanceolate. |
|
(70) |
+ |
Calyx teeth triangular to ovate-triangular. |
|
(71) |
|
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70 (69) |
Leaves ovate, 1.5-3.5 × 1-2.8 cm, margin crenate, adaxially dusty pubescent; petiole 5-13 mm |
|
33 Isodon lihsienensis |
+ |
Leaves lanceolate to oblanceolate, 0.8-1.5 × 0.5-0.7 cm, margin entire or obscurely few dentate above middle, adaxially densely tomentulose and papillate glandular; petiole 1-4 mm |
|
34 Isodon wikstroemioides |
|
|
|
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71 (69) |
Leaves 2.5-4.4 cm |
|
35 Isodon smithianus |
+ |
Leaves less than 2 cm. |
|
(72) |
|
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|
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72 (71) |
Leaves oblong-ovate, ovate, to broadly ovate, adaxially glandular pubescent, margin coarsely crenate or entire, apex rounded; cymes widely spaced |
|
36 Isodon parvifolius |
+ |
Leaves lanceolate to ovate-lanceolate, adaxially densely pilose, margin remotely crenulate-serrulate above middle to entire, apex acute; cymes compact, in terminal panicles |
|
37 Isodon tenuifolius |
|
|
|
|
73 (68) |
Cymes sessile or very short pedunculate; branchlets, inflorescences, and calyces densely hirtellous |
|
38 Isodon hirtellus |
+ |
Cymes conspicuously pedunculate; branchlets, inflorescences, and calyces not hirtellous. |
|
(74) |
|
|
|
|
74 (73) |
Leaves and branchlets densely tomentose when young, subglabrescent with age |
|
39 Isodon rubescens |
+ |
Leaves and branchlets hairy. |
|
(75) |
|
|
|
|
75 (74) |
Leaves ovate-lanceolate, 3.5-6(-10) × 1.5-2.5(-4) cm |
|
40 Isodon loxothyrsus |
+ |
Leaves ovate-oblong, rhomboid, to ovate-triangular, less than 4 cm. |
|
(76) |
|
|
|
|
76 (75) |
Branchlets and petioles antrorsely dull yellow velutinuous-pubescent; leaves ovate-oblong, 1-2.5 × 0.5-1.4 cm, base rounded |
|
41 Isodon xerophilus |
+ |
Branchlets and petioles gray tomentulose; leaves rhombic to triangular-ovate, 1.5-4 × 1-2.5 cm, base broadly cuneate to truncate |
|
42 Isodon rugosiformis |