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FOC | Family List | FOC Vol. 16 | Gentianaceae

3. Gentiana Sect. Kudoa (Masamune) Satake & Toyokuni ex Toyokuni, J. Jap. Bot. 35: 202. 1960.

多枝组 duo zhi zu

Kudoa Masamune, J. Soc. Trop. Agric. 2: 29. 1930; Gentiana sect. Monopodiae (Harry Smith) T. N. Ho; Gentiana subsect. Monopodiae Harry Smith.

Perennials. Roots linear-cylindric, lax, fleshy, arising from a collar. Stems branching monopodially. Stamens unequal (equal in Gentiana altorum). Seed coat alveolate, with simple shallow hexagonal pits.

Thirty-nine species: Asia; 30 species in China.


1 Base of vegetative stems forming a stout caudex; leaves (especially toward stem apex) and calyx lobes with white cartilaginous margin and midrib keel.   (2)
+ Base of vegetative axis usually slender; leaves and calyx lobes without white cartilaginous margin and midrib keel.   (4)
       
2 (1) Basal and rosette leaves distinctly contracted into petioles; calyx lobes obovate to spatulate, base distinctly narrowed   35 Gentiana stipitata
+ Basal and rosette leaves not distinctly contracted into petioles; calyx lobes lanceolate, elliptic, or elliptic-triangular, base scarcely narrowed.   (3)
       
3 (2) Corolla white with blue stripes and greenish spots when fresh, drying pale yellow or pale blue; lobes 5-6 mm   36 Gentiana szechenyii
+ Corolla ruddy purple with paler base when fresh, usually without stripes or spots; lobes 7-10 mm   37 Gentiana georgei
       
4 (1) Calyx lobes dissimilar in shape to upper leaves; plicae always asymmetrical.   (5)
+ Calyx lobes similar in shape to upper leaves; plicae always symmetrical.   (10)
       
5 (4) Flower in clusters of more than 3; stamens equal.   (6)
+ Flowers 1-3; stamens unequal.   (8)
       
6 (5) Basal rosettes none or poorly developed; lowermost stem leaves scalelike, membranous   29 Gentiana rigescens
+ Basal rosettes usually well developed; lowermost stem leaves not scalelike but with expanded herbaceous blades.   (7)
       
7 (6) Stems 3-15(-28) cm; corolla lobes caudate   30 Gentiana davidii
+ Stems (10-)20-50 cm; corolla lobes not caudate   31 Gentiana cephalantha
       
8 (5) Flowers 4merous; corolla nearly cylindric   32 Gentiana jingdongensis
+ Flowers 5merous; corolla funnelform.   (9)
       
9 (8) Blades of rosette leaves obovate to elliptic, abruptly narrowed into a petiole; corolla blue   33 Gentiana melandriifolia
+ Blades of rosette leaves narrowly spatulate to oblong-oblanceolate, gradually narrowed into a petiole; corolla rose   34 Gentiana duclouxii
       
10 (4) Stem leaves in whorls of 3-8.   (11)
+ Stem leaves opposite.   (17)
       
11 (10) Corolla lobes without a caudate tip.   (12)
+ Corolla lobes with a caudate tip to 3 mm.   (14)
       
12 (11) Margin of stem leaves and calyx lobes scabrous-ciliate   38 Gentiana ecaudata
+ Margin of stem leaves and calyx lobes smooth.   (13)
       
13 (12) Flowering stems with largest leaves toward apex; corolla 4.5-6 cm, lobe margin entire   39 Gentiana ternifolia
+ Flowering stems with largest leaves near middle; corolla 2.5-4.2 cm, lobe margin erose   40 Gentiana yakushimensis
       
14 (11) Style 5-7 mm; corolla narrowly obconic   44 Gentiana viatrix
+ Style 3-5 mm; corolla tubular to tubular-campanulate.   (15)
       
15 (14) Corolla lobes 2-2.5 mm, caudate tip less than 1 mm; stamens inserted just below middle of corolla tube   41 Gentiana tetraphylla
+ Corolla lobes 4-6 mm, caudate tip 2-2.5 mm; stamens inserted at basal part of corolla tube.   (16)
       
16 (15) Upper stem leaves and calyx lobes linear-spatulate, apex acute to obtuse   42 Gentiana hexaphylla
+ Upper stem leaves and calyx lobes linear, apex acuminate   43 Gentiana arethusae
       
17 (10) Corolla 1.5-2 cm; calyx lobes spreading   46 Gentiana altigena
+ Corolla 2.5-7.5 cm; calyx lobes straight or ascending.   (18)
       
18 (17) Basal rosettes or abbreviated vegetative shoots generally present at flowering time, their leaves usually conspicuously longer than those of flowering stems.   (19)
+ Basal rosettes and abbreviated vegetative shoots generally poorly developed or absent at flowering time, lowest leaves sometimes smaller and/or wider but not longer nor otherwise well differentiated.   (21)
       
19 (18) Stems papillate; corolla narrowly funnelform, 4-6 cm   50 Gentiana veitchiorum
+ Stems not papillate; corolla campanulate, 2.5-4.7 cm.   (20)
       
20 (19) Corolla lobes 4.5-6 mm; plicae 2-3 mm; anthers 3-3.5 mm   51 Gentiana nyalamensis
+ Corolla lobes 2.5-3.5 mm; plicae 0.5-1 mm; anthers 1.5-2 mm   52 Gentiana ornata
       
21 (18) Upper stem leaves orbicular, ovate, or elliptic.   (22)
+ Upper stem leaves linear or linear-lanceolate.   (25)
       
22 (21) Basal rosette absent; corolla cylindric   45 Gentiana prolata
+ Basal rosette present; corolla not cylindric.   (23)
       
23 (22) Corolla dark blue, obconic, lobes usually mucronate   47 Gentiana obconica
+ Corolla pale blue, not obconic, lobes not mucronate.   (24)
       
24 (23) All stem leaves elliptic   48 Gentiana altorum
+ Lower stem leaves ovate, upper ones elliptic   49 Gentiana caelestis
       
25 (21) Flowering stems 3-5 cm; corolla 3-3.8(-4.5) cm, lobes 3-4 mm; plicae 1.5-2 mm   55 Gentiana oreodoxa
+ Flowering stems 5-15(-20) cm; corolla 5-7.5 cm, lobes 6-9 mm; plicae 2-5 mm.   (26)
       
26 (25) Flowers usually pedicellate.   (27)
+ Flowers sessile.   (28)
       
27 (26) Calyx lobes and middle to upper leaves broadly linear, 2-3 mm wide; corolla campanulate-tubular, inflated at middle of tube; anthers 3.5-4 mm   53 Gentiana dolichocalyx
+ Calyx lobes and middle to upper leaves linear, often 1-1.5 mm wide; corolla narrowly obconic to funnelform; anthers 2.5-3 mm   54 Gentiana lawrencei
       
28 (26) Corolla 7-7.5 cm, inner surface blue-purple   56 Gentiana helophila
+ Corolla 5-6 cm, inner surface blue.   (29)
       
29 (28) Corolla conspicuously spotted; lower stem leaves oblong   57 Gentiana futtereri
+ Corolla unspotted or inconspicuously spotted; lower stem leaves linear-lanceolate   58 Gentiana sinoornata

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