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38. Leonurus Linnaeus, Sp. Pl. 2: 584. 1753.
益母草属 yi mu cao shu
Herbs annual, biennial, or perennial, erect. Leaves 3-7-lobed, basal ± palmately lobed, usually deciduous at anthesis; stem leaves entire, incised, or 3-lobed. Verticillasters many flowered, in long spikes; bracteoles subulate or spinescent, rigid or flaccid. Calyx obconical or tubular-campanulate, 5-veined, obscurely 2-lipped; upper lip straight, 3-toothed; lower lip 2-toothed, teeth coalescent, spreading or slightly spreading. Corolla white, reddish, to purplish, 2-lipped, tube exserted; upper lip oblong, obovate, or ovate-orbicular, margin entire, straight, villous or glabrous; lower lip straight or spreading, spotted, 3-lobed, lobes oblong-ovate or middle lobe shallowly cordate, larger than ovate lateral lobes. Stamens 4, anterior 2 longer, posterior 2 parallel; anther cells 2, parallel. Style apex equally 2-cleft, lobes subulate. Nutlets acutely triquetrous, base cuneate, apex truncate.
About 20 species: Africa, Asia, Europe, North America, South America; 12 species in China.
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1 |
Calyx indistinctly 2-lipped; anterior teeth not spreading; corolla tube not dilated, with or without distinct hairy annulus inside, if hairy annulus present then horizontal; upper lip of corolla oblong, base inconspicuously constricted; lower lip of corolla limb nearly straight; middle lobe much larger than lateral lobes, apex conspicuously emarginate, margin membranous; stamens nearly straight; leaves 3-lobed. |
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(2) |
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Calyx distinctly 2-lipped; anterior teeth strongly spreading; corolla tube distinctly oblique hairy annulate inside, conspicuously dilated above hairy annulus; upper lip of corolla obovate, base constricted; lower lip of corolla limb horizontally spreading, its 3 lobes subequal, oblong-ovate, margin revolute; anterior stamens projected, reflexed; leaves 5-7-lobed. |
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(7) |
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2 (1) |
Corolla less than 1 cm; without hairy annulus inside, puberulent |
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1 Leonurus chaituroides |
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Corolla more than 1 cm, distinctly ± horizontally scaly annulate inside. |
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(3) |
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3 (2) |
Leaves 3-lobed, lobes pinnately divided. |
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(4) |
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Leaves many lobed, lobes incised or coarsely serrate-dentate. |
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(5) |
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4 (3) |
Lobules of leaves more than 3 mm wide; floral leaves entire or rarely few dentate; corolla 1-1.2 cm; lower lip ca. as long as upper lip; calyx appressed puberulent |
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2 Leonurus japonicus |
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Lobules of leaves 1-3 mm wide, linear; floral leaves conspicuously 3-partite, lobules linear; corolla ca. 1.8 cm; lower lip ca. 3/4 as long as upper lip; calyx densely pilose especially at middle |
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3 Leonurus sibiricus |
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5 (3) |
Calyx teeth 5-10 mm; corolla 2.5-2.8 cm, reddish or reddish purple; leaves not corrugate, herbaceous |
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6 Leonurus macranthus |
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Calyx teeth 3-5 mm; corolla 1.5-2.1 cm, white, somewhat purple lineate; leaves corrugate, rigid. |
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(6) |
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6 (5) |
Plants densely villous |
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4 Leonurus villosissimus |
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Plants hirsute or strigose-hirtellous |
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5 Leonurus pseudomacranthus |
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7 (1) |
Inflorescences densely lanate-villous; bracteoles flaccid |
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12 Leonurus pseudopanzerioides |
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Inflorescences not lanate-villous; bracteoles rigid. |
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(8) |
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8 (7) |
Bracteoles and calyxes appressed pubescent, setose |
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8 Leonurus wutaishanicus |
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Bracteoles and calyxes appressed pubescent or appressed pubescent, white villous. |
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(9) |
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9 (8) |
Leaves 7-palmatilobed to 1/3 of their radius; lobes irregularly coarse dentate |
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7 Leonurus urticifolius |
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Leaves distinctly 5-palmatipartite from 2/3 to nearly entire radius; lobes pinnatilobulate, lobules linear to broadly lanceolate. |
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(10) |
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10 (9) |
Leaves numerous, lower stem leaves not early deciduous, lobes broadly cuneate, lobules broadly lanceolate; anterior teeth of calyx dilated in flower |
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10 Leonurus turkestanicus |
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Leaves few, lower stem leaves early deciduous, lobes narrowly cuneate to rhombic, lobules lanceolate to linear-lanceolate; anterior calyx teeth inconspicuously dilated in flower. |
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(11) |
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11 (10) |
Basal part of stems, nodes, rachises of inflorescences, and calyces appressed pubescent, subpatent white villous |
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11 Leonurus deminutus |
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Stems and calyxes appressed gray pubescent |
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9 Leonurus glaucescens |
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List of lower taxa
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