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FOC | Family List | FOC Vol. 2-3 | Thelypteridaceae

10. Leptogramma J. Smith, J. Bot. (Hooker). 4: 51. 1841.

茯蕨属 fu jue shu

Authors: Youxing Lin & Kunio Iwatsuki

Plants medium-sized, terrestrial. Rhizomes short and erect or ascending, with sparse scales; scales ovate-oblong or lanceolate, reddish brown, hairy. Fronds clustered; stipes deep stramineous, with sparse scales proximally, throughout with grayish white, mostly unicellular acicular long hairs and unicellular short setae; laminae oblong, hastate, or lanceolate, pinnate-pinnatifid; pinnae 7 or 8 pairs (or more), oblique or spreading, lanceolate, bases rounded or truncate, symmetrical, usually sessile, apices obtuse or acuminate; proximal 1 or 2 pairs or several pairs of pinnae free, distally ± adnate to rachises, proximal pair not or slightly shortened, sometimes prolonged, costae grooved adaxially, pinnae pinnatifid and usually reaching 1/2-2/3 of distance to costae; segments orbicular or oblong, entire and rounded at apices. Veinlets free, 3-6 pairs per segment, not forked, reaching margins, rarely reaching sinus. Laminae herbaceous or papery, when dry dark brown or brownish green, usually with acicular hairs or short setae on both surfaces, or with both mixed. Sori elongate, borne along veinlets, length slightly shorter than veinlets, exindusiate; sporangia each with 2-6 erect setae at top; spores bilateral, reniform, echinate on surfaces. x = 36.

About 15 species: tropical and subtropical areas of Asia, west to Africa; nine species (six endemic) in China.

"Leptogramma intermedia" (Ching in C. F. Zhang, Fl. Zhejiang 1: 154. 1993) was not validly published because no Latin description or diagnosis, or reference to such, was provided (Melbourne Code, Art. 39.1). It does not appear to represent a distinct species.

1 Proximal pair of pinnae longer than distal ones, hence laminae hastate.   1 L. tottoides
+ Proximal pair of pinnae similar length as distal ones or slightly shorter, laminae not hastate   (2)
2 (1) Proximal free pinnae with short stalks   (3)
+ All pinnae sessile   (4)
3 (2) Laminae not tapering to bases; free pinnae 5 pairs; sporangia each with 3 or 4 setae near top.   2 L. scallanii
+ Laminae slightly tapering to bases; free pinnae 9 pairs; sporangia each with 1 or 2 setae near top.   3 L. pozoi
4 (2) Stipes dark brown; laminae somewhat leathery and brown when dry; free pinnae 7-9 pairs.   4 L. huishuiensis
+ Stipes stramineous or deep stramineous; laminae herbaceous and greenish to deep green when dry (rarely dark brown); free pinnae 1-7 pairs   (5)
5 (4) Laminae dark brown when dry; pinnae without hairs on intercostal areas.   5 L. yahanensis
+ Laminae greenish to deep green when dry, with hairs on intercostal areas   (6)
6 (5) Rhizomes ascending; laminae yellowish green when dry.   6 L. jinfoshanensis
+ Rhizomes erect; laminae greenish to deep green when dry   (7)
7 (6) Laminae greenish when dry; sporangia subglabrous near tops.   7 L. centrochinensis
+ Laminae deep green when dry; sporangia each with 2-4 setae near top   (8)
8 (7) Pinnae with sparse acicular hairs adaxially, abaxially pubescent throughout.   8 L. himalaica
+ Pinnae with dense acicular hairs on both surfaces.   9 L. sinica

Lower Taxa


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