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FOC | Family List | FOC Vol. 17 | Lamiaceae

29. Leucas R. Brown, Prodr. 504. 1810.

绣球防风属 xiu qiu fang feng shu

Herbs or subshrubs. Leaves entire or dentate. Verticillasters few to many flowered, widely spaced, uniform or reduced in size at end of branches. Calyx tubular to obconical, rarely dilated, 10-veined, straight or curved, truncate or oblique at throat, projected on posterior side; teeth 8-10, usually equal in size. Corolla usually white, rarely yellow, purple, brownish, or scarlet, 2-lipped, tube included; upper lip straight, galeate, densely villous outside; lower lip longer, 3-lobed, middle lobe largest. Stamens ascending beneath upper lip of corolla, close together in pairs; anther cells 2, ovoid, divaricate, apex confluent, often orange. Style unequally 2-cleft at apex, posterior lobe short or obsolete. Nutlets ovoid, triquetrous, subtruncate at apex.

About 100 species: Africa, Asia, Australia, Pacific Islands, two naturalized in South America; eight species in China.

1 Verticillasters terminal, solitary; bracts lanceolate, overlapping to largely conceal flowers; calyx teeth very short   8 Leucas cephalotes
+ Verticillasters usually more than 1 per stem; bracts not concealing flowers; calyx teeth well developed.   (2)
2 (1) Calyx mouth not oblique.   (3)
+ Calyx mouth oblique, constricted or enlarged, reflexed or erect.   (5)
3 (2) Calyx densely villous inside throat; teeth spreading in fruit; leaves lanceolate to ovate-lanceolate, 6-9 × 1-3 cm; plants hirsute, densely covered with golden yellow hairs   3 Leucas ciliata
+ Calyx appressed silky to puberulent or subglabrous inside throat; teeth straight in fruit; leaves ovate; entire plant densely white tomentose.   (4)
4 (3) Leaves ovate-orbicular, 1-2.5(-3) × 0.8-1.5 cm, appressed silky to tomentose   1 Leucas chinensis
+ Leaves ovate, 2.5-4 cm × 1-2.5 cm, villous-tomentose (subglabrous in var. scaberula)   2 Leucas mollissima
5 (2) Leaves ovate to ovate-lanceolate, coarsely crenate-serrate; calyx mouth constricted, reflexed in fruit   6 Leucas martinicensis
+ Leaves linear, oblong, to oblong-lanceolate, entire or sparsely serrate; calyx mouth enlarged, straight.   (6)
6 (5) Plants slightly pubescent; calyx prominently oblique at mouth, teeth irregular or obsolete, apical tooth largest; leaves subentire to remotely undulate-serrate   7 Leucas lavandulifolia
+ Plants hispid; calyx slightly oblique at mouth, teeth regular, spinescent; leaves remotely serrate.   (7)
7 (6) Verticillasters ca. 1.5 cm in diam., few flowered; calyx tube not conspicuously ribbed adaxially   4 Leucas zeylanica
+ Verticillasters 2-2.5 cm in diam., many flowered; calyx tube conspicuously ribbed adaxially   5 Leucas aspera

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