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FOC | Family List | FOC Vol. 2-3 | Thelypteridaceae

5. Macrothelypteris (H. Itô) Ching, Acta Phytotax. Sin. 8: 308. 1963.

针毛蕨属 zhen mao jue shu

Authors: Youxing Lin & Kunio Iwatsuki

Thelypteris sect. Macrothelypteris H. Itô in Nakai & Honda, Nov. Fl. Jap. 4: 141. 1939.

Plants medium-sized, terrestrial, sometimes treelike, to 4 m tall. Rhizomes thick and short, erect, ascending or decumbent, with brown lanceolate long scales; scales thick, with acicular cilia along margins. Fronds clustered; stipes stramineous or reddish brown, glabrous, or with similar scales and after fallen with remaining lunate marks; laminae large, ovate-triangular, 3- or 4-pinnate-pinnatifid; pinnae and pinnules oblique or spreading and connected to each other by narrow wings along costae or costules; veins pinnate, free, lateral veins simple, sometimes forked. Laminae herbaceous or somewhat papery, yellowish green when dry, costae and costules rounded and raised adaxially, ± hairy on both sides and intercostal areas, rarely glabrous, hairs slender, grayish white acicular, unicellular or consisting of several cells, except above hairs along rachises, usually also with brown multicellular acicular thick hairs and few lanceolate or subulate thick scales and remaining protruding marks after scales fallen. Sori small, attached near ends of veinlets, exindusiate or with small and usually deciduous indusia; sporangia sometimes with shortly stalked capitate hairs near annuli. Spores bilateral, elliptic-reniform; perispores transparent and corrugate, echinate or minutely foveolate; exospore finely reticulate. x = 31.

About ten species: warmer parts of mainland Asia, NE Australia, Malesia, Pacific islands; seven species (one endemic) in China.

1 Rachises and costae covered with scales abaxially; scales inflated or incrassate at bases, leaving rough marks after falling   (2)
+ Rachises and costae without scales abaxially, or with grayish white hairs, or glabrous   (4)
2 (1) Plants 1-1.4 m tall; costae and costules with scales incrassate and not inflated at bases.   3 M. setigera
+ Plants 3-4 m tall, costae and costules covered with inflated bases of scales abaxially   (3)
3 (2) Scales on rachises denser, scales on costae entire.   1 M. ornata
+ Scales on rachises sparse, scales on costae with dense acicular hairs along margins.   2 M. polypodioides
4 (1) Pinnae glabrous or with unicellular acicular hairs   (5)
+ Pinnae with spreading multicellular acicular hairs abaxially   (6)
5 (4) Laminae herbaceous, when dry yellowish green; pinnules oblique distally and intersecting with costae at sharp angle; pinnae glabrous abaxially, at most with sparse acicular hairs along costae.   4 M. oligophlebia
+ Laminae thinly herbaceous, grass-green or dark green when dry; pinnules spreading and intersecting with costae at right angle; pinnae with more spreading acicular hairs abaxially.   5 M. viridifrons
6 (4) Laminae herbaceous, pinnules oblique distally and intersecting with costae at sharp angle; proximal pinnae with 2-2.5 cm stalks; hairs abaxially denser and elongate and 2.5-3 mm.   6 M. torresiana
+ Laminae thinly herbaceous, pinnules spreading and intersecting with costae at right angle; pinnae subsessile; hairs abaxially sparse and short and 1.5-2 mm.   7 M. contingens

Lower Taxa


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