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55. MADDENIA J. D. Hooker & Thomson, Hooker’s J. Bot. Kew Gard. Misc. 6: 381. 1854.

臭樱属 chou ying shu

Gu Cuizhi (Ku Tsue-chih); Bruce Bartholomew

Trees or shrubs, deciduous, polygamous dioecious. Branches unarmed. Winter buds large, oblong to ovoid, with several scales. Stipules large, persistent, margin glandular. Leaves alternate, simple; leaf blade margin with gland-tipped simple, double, or incised teeth. Inflorescences inserted apically on branchlets, racemose, many-flowered; bracts soon caducous. Pedicel short. Hypanthium campanulate. Sepals and petals essentially identical. Perianth segments 10–12, to 3 mm. Stamens 20–40, in 2 whorls, irregularly inserted on rim of hypanthium. Bisexual flowers: ovary superior, (1-or)2-loculed, glabrous; ovules 2 per locule, parallel, pendulous; style terminal, slender, ± as long as stamens; stigma disc-shaped. Female flowers: stamens reduced to staminodes; carpels 2; ovary glabrous; style much shorter than staminodes; stigma capitate. Fruit a 1-or 2-seeded drupe, oblong, ± flat; mesocarp thin, fleshy, not splitting when ripe; endocarp bony, ovoid, 3-angled, apex acute.

About seven species: China, Himalayas; six species (five endemic) in China.


1 Leaf blade abaxially glabrous; branchlets glabrous or pubescent.   (2)
+ Leaf blade abaxially densely pubescent at least on veins; branchlets densely pubescent.   (4)
       
2 (1) Leaf blade abaxially whitish, glaucous, margin irregularly or doubly serrate; secondary veins 14–18 on each side of midvein.   1 M. hypoleuca
+ Leaf blade abaxially green, not glaucous, margin doubly serrate or incisedly doubly serrate; secondary veins 9–15 on each side of midvein.   (3)
       
3 (2) Branchlets densely pubescent; leaf blade 5–10(–15) × 3–5(–8) cm, margin incisedly double serrate.   2 M. incisoserrata
+ Branchlets sparsely pubescent; leaf blade 4–5 × 2–2.5 cm, margin not incisedly doubly serrate.   3 M. fujianensis
       
4 (1) Branchlets of first year’s growth villous; winter buds purplish red; hypanthium villous; style ± as long as stamens.   6 M. himalaica
+ Branchlets of first year’s growth pubescent or tomentose-pubescent; winter buds purplish brown; hypanthium pubescent; style longer than stamens.   (5)
       
5 (4) Winter bud scales to 2 × 1 cm; stipules linear to lanceolate, herbaceous; inflorescence bracts lanceolate to ovate-lanceolate.   4 M. hypoxantha
+ Winter bud scales to 1.5 × 0.8 cm; stipules lorate-lanceolate, membranous; inflorescence bracts oblong.   5 M. wilsonii

Lower Taxa


 

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