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23. Mazus Loureiro, Fl. Cochinch. 2: 385. 1790.

通泉草属 tong quan cao shu

Hornemannia Willdenow.

Herbs, relatively small. Stems terete or rarely quadrangular (Mazus lanceifolius), erect or procumbent and rooting from lower nodes. Leaves in a rosette or opposite, often upper leaves alternate; petiole winged. Racemes secund; bracts small. Bracteoles present or absent. Flowers small. Calyx funnelform or campanulate, 5-lobed. Corolla 2-lipped, palate with 2 longitudinal plaits; lower lip 3-lobed; upper lip 2-lobed. Stamens 4, didynamous, inserted on corolla tube; anther locules divergent, apically connivent. Ovary hairy or glabrous. Style glabrous; stigma 2-lamellate. Capsule compressed, included in cupular persistent calyx, loculicidal. Seeds small, numerous.

About 35 species: China, India, Indonesia, Japan, Korea, Malaysia, Mongolia, Philippines, Russia, Vietnam; Australia, New Zealand; 25 species in China.


1 Stems quadrangular   25 Mazus lanceifolius
+ Stems terete or somewhat ribbed, never quadrangular.   (2)
       
2 (1) Ovary hairy; stems basally woody with age; calyx veins conspicuous.   (3)
+ Ovary glabrous; stems herbaceous (woody in M. celsioides); calyx veins inconspicuous.   (6)
       
3 (2) Plants relatively stout, erect, never rooting from nodes; flowers ca. 1.5 cm or more; calyx funnelform, 0.8-1.6 cm in fruit, over 1 cm in diam.   (4)
+ Plants slender, procumbent, rooting from nodes; flowers less than 1.5 cm; calyx campanulate, 0.3-0.8 cm in fruit, less than 1 cm in diam.   (5)
       
4 (3) Stem leaves sessile; corolla 1.5-2 cm; capsule ovoid   1 Mazus stachydifolius
+ Stem leaves petiolate; corolla ca. 2.6 cm; capsule globose   2 Mazus caducifer
       
5 (3) Basal leaves caudate; calyx lobes triangular-lanceolate; corolla upper lip lobes apically acute, margin entire   3 Mazus spicatus
+ Basal leaves not caudate; calyx lobes ovate; corolla upper lip lobes apically subtruncate, margin with few small irregular teeth near apex   4 Mazus xiuningensis
       
6 (2) Plants stoloniferous.   (7)
+ Plants not stoloniferous.   (15)
       
7 (6) Plants glabrous or soon glabrescent.   (8)
+ Plants pubescent, sometimes only when young.   (11)
       
8 (7) Stems less than 10 cm tall.   (9)
+ Stems more than 10 cm tall.   (10)
       
9 (8) Basal leaves thin papery; bracts ca. 1 mm   7 Mazus rockii
+ Basal leaves fleshy; bracts 5-12 mm   15 Mazus longipes
       
10 (8) Stems erect to obliquely ascending, 10-15 cm tall; middle lobe of corolla lower lip obovate, smaller than lateral lobes   21 Mazus miquelii
+ Stems creeping, to 30 cm; middle lobe of corolla lower lip narrowly ovate, longer than lateral lobes   23 Mazus gracilis
       
11 (7) Stem leaves often alternate.   (12)
+ Stem leaves often opposite or plants scapose.   (13)
       
12 (11) Stems erect to obliquely ascending, 10-15 cm tall   21 Mazus miquelii
+ Stems procumbent, 30-45 cm   24 Mazus procumbens
       
13 (11) Stems less than 8 cm tall   16 Mazus surculosus
+ Stems to 30 cm tall.   (14)
       
14 (13) Basal leaf blade suborbicular to broadly ovate; pedicel 1.5-2.5 cm; calyx lobes less than 1/3 of calyx length   20 Mazus henryi
+ Basal leaf blade spatulate; pedicel 0.5-0.6 cm; calyx lobes ca. 1/2 of calyx length   22 Mazus alpinus
       
15 (6) Stem leaves 2-4 pairs or numerous.   (16)
+ Stem leaves absent or 1 or 2.   (21)
       
16 (15) Calyx lobes ca. 1/3 of calyx length.   (17)
+ Calyx lobes ca. 1/2 of calyx length.   (18)
       
17 (16) Stems basally woody; leaf margin irregularly serrate, pandurately lobed below middle; stems and leaves abaxially green   12 Mazus celsioides
+ Stems herbaceous; leaves margin irregularly sinuate-toothed; stems and leaves abaxially purple   13 Mazus solanifolius
       
18 (16) Perennials, white villous.   (19)
+ Annuals, glabrous or sparsely pubescent.   (20)
       
19 (18) Stem leaves crowded; corolla purple-red, lower middle lobe longer than other lobes   14 Mazus kweichowensis
+ Stem leaves 2-4 pairs; corolla blue-purple, lower middle lobe shorter than other lobes   18 Mazus saltuarius
       
20 (18) Pedicel 0.3-1.2 cm; corolla lower lip lobes obovate; upper lobes ovate-triangular   17 Mazus pumilus
+ Pedicel ca. 1.5 cm; corolla lower lip lobes broadly ovate; upper lobes oblong-ovate   19 Mazus fukienensis
       
21 (15) Corolla 2-3 cm, lobes usually erose or fimbriate apically; leaves usually to 20 cm.   (22)
+ Corolla less than 2 cm, lobes not fimbriate; leaves less than 12 cm.   (23)
       
22 (21) Plants glabrous or sparsely villous; leaves thick papery to subleathery; pedicel 1-15 cm   5 Mazus omeiensis
+ Plants densely white or rusty pubescent; leaves papery; pedicel to 4 cm   6 Mazus pulchellus
       
23 (21) Plants mostly over 10 cm; flowers usually over 10.   (24)
+ Plants less than 10 cm; flowers less than 10.   (25)
       
24 (23) Pedicel ca. 5 cm; leaves oblanceolate, margin sharply toothed; flowering stems erect   10 Mazus lecomtei
+ Pedicel 1-3 cm; leaves obovate-spatulate, margin obtusely toothed or pinnately lobed; flowering stems not erect   11 Mazus fauriei
       
25 (23) Leaf margin entire; flowers 1-3; pedicel to 3 cm   9 Mazus oliganthus
+ Leaf margin dentate or shallowly lobed; flowers 4-10; pedicel less than 2 cm.   (26)
       
26 (25) Plants glabrous or subglabrous; calyx lobes subulate-triangular; corolla yellow or dark purple   7 Mazus rockii
+ Plants sparsely white villous; calyx lobes oblong-lanceolate; corolla white or white with purple spots   8 Mazus humilis

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