|
2. Meliosma Blume, Catalogus. 10. 1823.
泡花树属 pao hua shu shu
Millingtonia Roxburgh, Pl. Coromandel 3: 50. 1820 ["1819"], not Linnaeus f. (1782); Wellingtonia Meisner (1840), not Lindley (1853).
Trees or shrubs, evergreen or deciduous. Buds naked, brownish tomentose. Leaves simple or odd pinnate, leaves or leaflets entire or often dentate; petiole usually thickened at base. Inflorescence terminal, sometimes axillary, a pyramidal panicle, usually profusely flowered but sometimes sparsely so, up to 4 × branched and very large. Flowers numerous, 1-3 mm in diam., hermaphroditic, zygomorphic, sessile or shortly pedicellate. Sepals [3 or](4 or)5(-9), imbricate, with bracts beneath. Petals 5, imbricate, unequal, 3 outer ones larger, usually suborbicular or reniform, convex, 2 inner ones much smaller, bifid or entire, sometimes 3-lobed, ± adherent to base of filaments of fertile stamens, concealed under outer petals at bud stage. Fertile stamens 2, opposite 2 inner petals; filaments flattened, short, incurved at apex into a cup-shaped connective; anthers 2-celled, globose or elliptic; staminodes 3, opposite outer sepals, adherent to their bases. Disk cup-shaped or shallowly cup-shaped, 5-dentate. Ovary sessile, 2(or 3)-loculed, with 1 or 2 ovules per locule. Fruit a drupe, small; mesocarp fleshy; endocarp stony or crustaceous. Seed ± globose, usually somewhat concave ventrally, without endosperm.
About 50 species: SE Asia, Central and South America; 29 species (ten endemic) in China.
The reader may also wish to refer to Beusekom, Revision of Meliosma (Sabiaceae), section Lorenzanea excepted, living and fossil, geography and phylogeny (Blumea 19: 355-529. 1971). Meliosma cavaleriei H. Léveillé (Repert. Spec. Nov. Regni Veg. 9: 457. 1911) is probably a synonym of Ampelopsis chaffanjonii (H. Léveillé & Vaniot) Rehder in the Vitaceae (see p. 183).
|
|
|
|
| 1 |
Leaves pinnate |
|
(2) |
| + |
Leaves simple |
|
(10) |
| |
|
|
|
| 2 (1) |
Terminal leaflet (rarely 2 leaflets) with articulation on petiolule; sepals usually 4; outer petals broadly reniform or other shapes, slightly broader than long |
|
(3) |
| + |
Terminal leaflets 3, petiolules not articulate; sepals 5; outer petals suborbicular or broadly elliptic, narrower than long |
|
(4) |
| |
|
|
|
| 3 (2) |
Leaflets abaxially crinite at vein axils; panicle 12-30 cm; 2 inner petals acutely bifid |
|
28 M. alba |
| + |
Leaflets abaxially not crinite at vein axils; panicle 40-45(-60) cm; 2 inner petals obtusely bifid |
|
29 M. veitchiorum |
| |
|
|
|
| 4 (2) |
Leaflets leathery, abaxially pubescent or glabrous |
|
(5) |
| + |
Leaflets subleathery, abaxially pubescent, lanuginous, or glandulous (except in M. pinnata) |
|
(6) |
| |
|
|
|
| 5 (4) |
Leaflets narrowly elliptic, elliptic-lanceolate, or lanceolate, margin entire or rarely 1- or 2-toothed near apex, only sparsely pubescent on midvein. |
|
21 M. angustifolia |
| + |
Leaflets narrowly ovate, ovate-lanceolate, or oblong-elliptic, margin sparsely spinose-serrate, both surfaces glabrous, or only crinite at axils of lateral veins. |
|
22 M. rhoifolia |
| |
|
|
|
| 6 (4) |
Leaflets basally usually cuneate, glabrous on both surfaces; petiole terete. |
|
23 M. pinnata |
| + |
Leaflets basally usually obtuse-rounded, ± pubescent; petiole grooved |
|
(7) |
| |
|
|
|
| 7 (6) |
Leaflets adaxially shortly pubescent, abaxially pubescent or subglabrous. |
|
27 M. oldhamii |
| + |
Leaflets adaxially glabrous except pubescent on midvein and lateral veins |
|
(8) |
| |
|
|
|
| 8 (7) |
Leaflets abaxially pale, claviform-glandular, but crinite at vein axils. |
|
26 M. glandulosa |
| + |
Leaflets abaxially light green, sparsely pilose or lanuginous |
|
(9) |
| |
|
|
|
| 9 (8) |
Branchlets brown; leaflets ovate or ovate-lanceolate, abaxially and on inflorescences pilose or lanuginous. |
|
24 M. arnottiana |
| + |
Branchlets red; leaflets narrowly oblong, narrowly ovate, or narrowly obovate-elliptic, abaxially and on inflorescences sparsely pubescent. |
|
25 M. kirkii |
| |
|
|
|
| 10 (1) |
Lateral veins straight or nearly so |
|
(11) |
| + |
Lateral veins distinctly ascending |
|
(15) |
| |
|
|
|
| 11 (10) |
Two inner petals entire, longer than fertile stamens. |
|
5 M. myriantha |
| + |
Two inner petals bifid, or sometimes between the 2 lobes, with a smaller one, shorter than fertile stamens |
|
(12) |
| |
|
|
|
| 12 (11) |
Panicle pendulous, rachis and branches distinctly zigzag. |
|
4 M. flexuosa |
| + |
Panicle erect, lateral branches not pendulous, rachis and branches straight or slightly flexuose |
|
(13) |
| |
|
|
|
| 13 (12) |
Leaf margin sharply biserrate, abaxially curved pilose, inconspicuously crinite at vein axils. |
|
1 M. dilleniifolia |
| + |
Leaf margin serrate, or seldomly 1- or 2-biserrate, with straight hairs or sparsely pubescent, conspicuously crinite at vein axils |
|
(14) |
| |
|
|
|
| 14 (13) |
Leaf blade obovate-cuneate or narrowly obovate-cuneate, apex shortly acuminate, margin serrate. |
|
2 M. cuneifolia |
| + |
Leaf blade obovate, apex subtruncate, with short acute tip, margin undulate. |
|
3 M. parviflora |
| |
|
|
|
| 15 (10) |
Panicle narrow, 4-7 cm wide, lateral branches short, less than 1 cm in diam |
|
(16) |
| + |
Panicle broad, more than 8 cm wide |
|
(17) |
| |
|
|
|
| 16 (15) |
Petiole narrowly winged, adaxially glabrous; 2 inner petals 2-lobed. |
|
6 M. paupera |
| + |
Petiole densely pubescent, adaxially scattered pubescent; 2 inner petals bifid to half. |
|
7 M. bifida |
| |
|
|
|
| 17 (15) |
Leaves 15-40 × 4-16 cm, lateral veins 15-28 pairs |
|
(18) |
| + |
Leaves usually shorter than 15 cm, if to 15 cm, less than 5 cm wide, lateral veins fewer than 15 pairs |
|
(21) |
| |
|
|
|
| 18 (17) |
Leaf blade abaxially densely lanuginous; sepals 5, outside with 4 or 5 equal bracts. |
|
8 M. thomsonii |
| + |
Leaf blade glabrous or sparsely pubescent; sepals 4 or 5, without equal bracts |
|
(19) |
| |
|
|
|
| 19 (18) |
Ovary densely pubescent, drupes persistently pubescent; 2 inner petals entire. |
|
9 M. thorelii |
| + |
Ovary and drupes pubescent; 2 inner petals bifid |
|
(20) |
| |
|
|
|
| 20 (19) |
Petiole 1.5-2.5 cm; branchlets pubescent; drupes globose. |
|
10 M. simplicifolia |
| + |
Petiole 3-5 cm; branchlets glabrous; drupes obovoid or subglobose. |
|
11 M. longipes |
| |
|
|
|
| 21 (17) |
Leaf margin serrate |
|
(22) |
| + |
Leaf margin entire, rarely 1- or 2-toothed near apex |
|
(24) |
| |
|
|
|
| 22 (21) |
Abaxial surface of leaves and inflorescences sparsely pilose or densely tomentose. |
|
18 M. rigida |
| + |
Abaxial surface of leaves and inflorescences sparsely procumbent pubescent |
|
(23) |
| |
|
|
|
| 23 (22) |
Petiole densely lanuginous; panicle 2 or 3 × branched. |
|
20 M. yunnanensis |
| + |
Petiole sparsely pubescent; panicle 3 or 4 × branched. |
|
19 M. fordii |
| |
|
|
|
| 24 (21) |
Petiole slender, 2.5-6.5(-10) cm, base terete, swollen; leaves clustered terminally |
|
(25) |
| + |
Petiole stouter, usually shorter than 2.5 cm, base not swollen; leaves scattered on branches |
|
(26) |
| |
|
|
|
| 25 (24) |
Leaves abaxially pale, densely yellowish brown squamellate. |
|
12 M. squamulata |
| + |
Leaves abaxially light brown, sparsely pilose. |
|
13 M. dumicola |
| |
|
|
|
| 26 (24) |
Leaves abaxially glabrous or only crinite at vein axils |
|
(27) |
| + |
Leaves densely ferruginous lanuginous or pilose |
|
(28) |
| |
|
|
|
| 27 (26) |
Leaves leathery, abaxially glabrous; flowers ca. 2 mm in diam., 2 inner petals entire. |
|
14 M. henryi |
| + |
Leaves papery, crinite at vein axils; flowers 1-1.5 mm in diam., 2 inner petals bifid. |
|
15 M. callicarpifolia |
| |
|
|
|
| 28 (26) |
Leaves leathery, adaxially glabrous except on midvein and lateral veins; 2 inner petals bifid to half. |
|
16 M. laui |
| + |
Leaves membranous or papery, adaxially pubescent, at least sparsely so; 2 inner petals 2-lobed. |
|
17 M. velutina |
|
|
|
List of lower taxa
| Related Links (opens in a new window) |
|
Treatments in Other Floras @ www.efloras.org
Other Databases
|
|
|
|
|
|
|
