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FOC | Family List | FOC Vol. 19 | Rubiaceae

57. Mycetia Reinwardt, Syll. Pl. Nov. 2: 9. 1825.

腺萼木属 xian e mu shu

Authors: Tao Chen & Charlotte M. Taylor

Adenosacme Wallich ex Miquel, nom. illeg. superfl.

Small shrubs, unarmed, usually rather succulent; bark often straw-yellow to nearly white, soft, often corky. Raphides present. Leaves opposite, sometimes weakly to markedly anisophyllous, frequently somewhat asymmetrical or falcate, apparently without domatia, often with margins crisped; tertiary venation often closely reticulate and prominulous below and frequently also above; stipules persistent with leaves or caducous, interpetiolar, triangular or sometimes leaflike, often well developed, sometimes with glands. Inflorescences terminal, pseudoaxillary, cauline, or perhaps sometimes axillary, cymose to paniculiform or subcapitate, several to many flowered, pedunculate, bracteate or bracts reduced; bracts sometimes leaflike, sometimes glandular. Flowers pedicellate, bisexual, at least usually distylous. Calyx limb deeply (4 or)5(or 6)-lobed; lobes frequently somewhat unequal, usually with sessile to stalked marginal glands. Corolla yellow or white, tubular, funnelform, salverform, or campanulate, sometimes gibbous at base, inside glabrous or usually pubescent; lobes (4 or)5(or 6), in bud valvate-induplicate. Stamens (4 or)5(or 6), inserted in throat or above middle of corolla tube in short-styled flowers, inserted near base of corolla tube in long-styled flowers; filaments short or reduced; anthers apparently dorsifixed, included or partially exserted. Ovary 2(-5)-celled, ovules numerous in each cell on fleshy axile placentas; stigmas 2(-5), included in short-styled flowers or exserted in long-styled flowers. Fruit white or perhaps brown, baccate or perhaps capsular and irregularly dehiscent, leathery to fleshy or spongy, subglobose, with calyx limb persistent; seeds numerous, small, angled, with testa somewhat granular.

About 45 species: tropical and subtropical Asia; 15 species (ten endemic) in China.

This genus was reviewed for Thailand by Fukuoka (Acta Phytotax. Geobot. 40: 107-118. 1989), for the Indian subcontinent by Deb (Bull. Bot. Surv. India 28(1-4): 114-132. 1986), and for China by H. S. Lo (Guihaia 11: 105-116. 1991). Mycetia is said to belong to Isertieae but also, anomalously, to have raphides. H. S. Lo (in FRPS 71(1): 314. 1999) reported that the inflorescences may sometimes be axillary, but this has not been noted by other authors; this description may be using the term "axillary" for the position elsewhere considered "pseudoaxillary." Lo also reported that the anthers may sometimes be partially exserted, which has not been reported by other authors nor seen on specimens studied by us.

1 Pedicels mostly or all 9-20 mm   (2)
+ Flowers subsessile or with pedicels mostly or all 8 mm or shorter   (5)
2 (1) At least some nodes with markedly anisophyllous leaves, larger leaf 3-10 × (or more) as long as smaller leaf; calyx lobes 2.5-3 mm; corolla tube 12-16 mm.   7 M. gracilis
+ Leaves generally isophyllous, larger leaf at most 2 × as long as smaller leaf; calyx lobes 0.8-6 mm; corolla tube 5-10 mm (unknown in M. yunnanica)   (3)
3 (2) Calyx lobes 3-6 mm, markedly longer than length of hypanthium together with unlobed basal part of limb.   2 M. bracteata
+ Calyx lobes 0.8-3 mm, shorter than or ± equal to length of hypanthium together with unlobed basal part of limb   (4)
4 (3) Inflorescences terminal and pseudoaxillary; calyx lobes 0.8-1 mm.   4 M. brevisepala
+ Inflorescences pseudoaxillary or possibly axillary at lower stem nodes, often below leaves; calyx lobes 2.5-3 mm.   15 M. yunnanica
5 (1) At least some calyx limbs and sometimes also bracts bearing few to numerous marginal glands on short to well-developed flexuous stipes   (6)
+ Calyx limbs and bracts without raised or stipitate marginal glands   (9)
6 (5) Stipules elliptic-oblong, obovate, or suborbicular and narrowed to stipitate at base, at base less than 1/2 as wide as maximum width; calyx lobes with 1-3 pairs of glands.   14 M. sinensis
+ Stipules lanceolate, oblong-lanceolate, or ovate, at base straight or narrowed but more than 2/3 as wide as maximum width; calyx lobes with numerous stipitate glands   (7)
7 (6) Corolla with tube 4-6 mm; leaves moderately to densely hispidulous or hirtellous adaxially.   9 M. hirta
+ Corolla with tube 7-14 mm; leaves glabrous or sparsely strigillose or hispidulous adaxially   (8)
8 (7) Petioles 1-15 mm; calyx lobes 1-1.5 mm; corolla tube 7-10 mm.   6 M. glandulosa
+ Petioles 6-60 mm; calyx lobes 1.5-4 mm; corolla tube 10-14 mm.   11 M. longifolia
9 (5) Calyx lobes 1.5-2 mm; corolla tube 5-7 mm (mature corollas unknown in M. hainanensis)   (10)
+ Calyx lobes 3-6 mm; corolla tube 7-17 mm (corolla unknown in M. macrocarpa)   (11)
10 (9) Calyx densely hirtellous; flowers sessile or subsessile; stipules ovate, not strongly contracted at base.   8 M. hainanensis
+ Calyx glabrous; flowers pedicellate; stipules elliptic-oblong, obovate, or suborbicular, contracted to stipitate at base.   14 M. sinensis
11 (9) Stipules broadly elliptic, subovate, or suborbicular, 5-10 mm   (12)
+ Stipules triangular, lanceolate, suborbicular, or ovate, 3-6 mm   (13)
12 (11) Leaves glabrous on both surfaces; fruit ca. 6 mm; Yunnan.   12 M. macrocarpa
+ Leaves strigillose to hirtellous abaxially; fruit 7-8 mm; Xizang.   13 M. nepalensis
13 (11) Corolla tube 7-8 mm; leaves subsessile or with petioles up to 15 mm   (14)
+ Corolla tube 10-17 mm; leaves with petioles 10-70 mm   (15)
14 (13) Leaves subsessile; inflorescences several flowered (i.e., with 5-7 flowers).   3 M. brevipes
+ Leaves with petioles 3-15 mm; inflorescences many flowered (i.e., with 9-40 flowers).   5 M. coriacea
15 (13) Flowers sessile or subsessile; petioles 10-20 mm.   1 M. anlongensis
+ Flowers pedicellate with pedicels 2.5-7 mm; petioles 20-70 mm.   10 M. longiflora

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