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FOC | Family List | FOC Vol. 4 | Podocarpaceae

3. Nageia Gaertner, Fruct. Sem. Pl. 1: 191. 1788.

竹柏属 zhu bai shu

Trees evergreen, dioecious or rarely monoecious; crown columnar. Leaves spirally arranged or in decussate, opposite pairs on leading shoots, opposite or subopposite on lateral shoots, ± monomorphic, adult leaves similar to juvenile leaves but often larger or wider (although juvenile leaves larger in Nageia wallichiana), more than 5 mm; petiole twisted through 90° ; blade broadly ovate-elliptic to oblong-lanceolate, without obvious midvein but with many, slender, parallel, longitudinal veins converging toward base and apex, stomatal lines abaxial or rarely on all surfaces. Pollen cones axillary, solitary or clustered in small, spikelike groups of 3-6, borne on naked peduncles, ovoid-cylindric, with basal sterile scales; pollen 2-saccate. Seed-bearing structures terminal on short, scaly, axillary branchlets, solitary or occasionally paired; bracts usually obsolete, scarcely thicker than peduncle, rarely succulent and thicker than peduncle; ovule inverted. Epimatium wholly enveloping seed, leathery, with bluish black bloom when ripe. Seed drupelike, globose.

Five to seven species: Bangladesh, Cambodia, China, India, Indonesia, Japan (including Ryukyu Islands), Laos, Malaysia, Myanmar, Philippines, Thailand, Vietnam; three species in China.

The leaves of Nageia strongly differ from those of Podocarpus in their numerous, parallel veins and absence of a midvein, and are superficially much more similar to those of Agathis (Araucariaceae). The Chinese species of Nageia were treated in FRPS under Podocarpus. D. Z. Fu (Acta Phytotax. Sin. 30: 515-528. 1991) placed the genus in its own family, Nageiaceae, but this view has since been refuted by several workers using different lines of evidence.

1 Leaf blade with stomatal lines present on both surfaces; receptacle thick, very succulent   1 Nageia wallichiana
+ Leaf blade with stomatal lines present on abaxial surface only, sometimes scarcely visible; receptacle obsolete or absent.   (2)
2 (1) Leaf blade usually 8-18 cm; receptacle absent; seed 1.5-1.8 cm in diam.   2 Nageia fleuryi
+ Leaf blade to 9 cm; receptacle obsolete; seed 1-1.5 cm in diam.   3 Nageia nagi

Lower Taxa


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