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FOC | Family List | FOC Vol. 11 | Rutaceae

10. Phellodendron Ruprecht, Bull. Cl. Phys.-Math. Acad. Imp. Sci. Saint-Pétersbourg. 15: 353. 1857.

黄檗属 huang bo shu

Authors: Prof. Dianxiang Zhang & Thomas G. Hartley

Trees, deciduous, dioecious. Bark corky, fissured and gray or grayish brown at maturity; inner bark (phloem) yellow. Branchlets dark purplish. Leaves opposite, odd-pinnate; base of petiole excavated, concealing axillary bud; leaflet blades with oil glands restricted to margin, margin subentire to minutely crenulate or serrulate. Inflorescences terminal, thyrsiform. Sepals 5(-8), connate at base. Petals 5(-8), narrowly imbricate in bud. Stamens 5(-7), distinct. Disk columnar. Male flowers: stamens to 1.5 × as long as petals; gynoecium rudimentary, of 5(or 6) fingerlike carpels, basally connate, apically ± divergent. Female flowers: stamens rudimentary, ligulate, much shorter than petals; gynoecium 5(-10)-loculed, syncarpous; ovules 1 per locule; stigma peltate. Fruit a drupaceous berry, black or purplish black, 5(-10)-loculed; exocarp fleshy; mesocarp undifferentiated; endocarp thinly cartilaginous. Seeds brown or black, asymmetrically ellipsoid, dull, inconspicuously rugulose; seed coat with inner layer of dense black sclerenchyma and outer layer of compact parenchymatous tissue; endosperm ± scant; embryo straight; cotyledons elliptic, flattened; hypocotyl superior.

Two to four species: E and SE Asia; two species (one endemic) in China.

1 Inflorescences and infructescences ± lax, rachis, branches, and pedicels slender; leaflet blades papery to thinly papery.   1 P. amurense
+ Inflorescences and infructescences ± compact, rachis, branches, and pedicels robust; leaflet blades papery.   2 P. chinense

Lower Taxa


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