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FOC | Family List | FOC Vol. 7 | Lauraceae

12. Phoebe Nees, Syst. Laur. 98. 1836.

楠属 nan shu

Authors: Fa-Nan Wei & Henk van der Werff

Evergreen trees or shrubs. Leaves alternate, pinnately veined. Flowers bisexual, cymose paniculate or subracemose. Perianth lobes 6, equal in size or sometimes outer ones slightly short, becoming leathery or woody after flowering. Fertile stamens 9, in 3 series; 1st and 2nd series without glands and with introrse 4-celled anthers; 3rd series with 2 glands and extrorse 4-celled anthers. Staminodes triangular or sagittate. Ovary ovoid or globose; stigma dish-shaped or capitate. Fruit ovoid, ellipsoid, or globose, rarely oblong, base surrounded by persistent and enlarged perianth lobes; fruiting pedicel not thickened or conspicuously thickened.

Up to 100 species: tropical and subtropical Asia; 35 species (27 endemic) in China.


1 Perianth lobes outside and inflorescences glabrous or appressed puberulent   (2)
+ Perianth lobes outside and inflorescences densely pubescent, villous, or tomentose   (15)
       
2 (1) Midrib of leaf blade completely elevated adaxially   (3)
+ Midrib of leaf blade partly or completely impressed adaxially   (8)
       
3 (2) Branchlets and leaf blade abaxially glaucous.   1 P. lichuanensis
+ Branchlets and leaf blade not as above   (4)
       
4 (3) Fruit oblong or ellipsoid   (5)
+ Fruit ovoid   (6)
       
5 (4) Leaf blade elliptic, 7-13(-15) × 2-4 cm, lateral veins 6 or 7 pairs; petiole 1-2 cm; infructescences 3-7 cm; fruit ellipsoid, 1.1-1.3 cm × 5-7 mm.   5 P. yaiensis
+ Leaf blade broadly oblong-lanceolate or oblong-oblanceolate, 10-28 × 4-8 cm, lateral veins 9-15 pairs; petiole 2-4 cm; infructescences 10-17 cm; fruit oblong, 1.6-1.8 cm × ca. 8 mm.   6 P. hainanensis
       
6 (4) Leaf blade narrowly lanceolate, usually 15-25 × 1-2.5 cm.   2 P. angustifolia
+ Leaf blade lanceolate, elliptic, or elliptic-lanceolate, much wider, 3-6.5 cm in diam   (7)
       
7 (6) Leaf blade thickly papery, veinlets not foveolate on both surfaces; inflorescences slender; pedicel glaucous; fruit beaked at apex.   3 P. lanceolata
+ Leaf blade leathery, veinlets foveolate on both surfaces; inflorescences thick; pedicel not glaucous; fruit not beaked at apex.   4 P. calcarea
       
8 (2) Inflorescences mostly slender, much branched, each branch with persistent bracts at base; leaf blade lateral veins and veinlets deeply impressed adaxially.   7 P. kwangsiensis
+ Inflorescences thick, each branch without persistent bracts at base; leaf blade lateral veins and veinlets not impressed or slightly impressed adaxially   (9)
       
9 (8) Fruit globose or subglobose   (10)
+ Fruit ovoid   (13)
       
10 (9) Flowers large, 5-6 mm; petiole to 4 cm, 2-3 mm thick.   8 P. chinensis
+ Flowers smaller, 2.5-3.5 mm; petiole not more than 2.5 cm, more slender   (11)
       
11 (10) Leaf blade not lustrous on both surfaces, abaxially not glaucescent.   11 P. minutiflora
+ Leaf blade lustrous on both surfaces, abaxially glaucescent   (12)
       
12 (11) Leaf blade leathery, ovate-elliptic or lanceolate, 4-6 × 1-3.5 cm, lateral veins 5-7 pairs.   9 P. microphylla
+ Leaf blade thickly leathery, oblong-lanceolate or elliptic, 7-12(-15) × 2-4.5 cm, lateral veins 12-15 pairs.   10 P. faberi
       
13 (9) Young leaf blade densely appressed sericeous abaxially; perianth lobes conspicuously ciliate.   12 P. hunanensis
+ Young leaf blade glabrous or sparsely pubescent abaxially; perianth lobes glabrous or somewhat hairy at margins   (14)
       
14 (13) Leaf blade elliptic, lateral veins 6-9 pairs, inconspicuous on both surfaces, margin revolute; perianth lobes glabrate inside.   13 P. nigrifolia
+ Leaf blade oblanceolate, lanceolate, pandurate-oblanceolate, or pandurate-obovate, lateral veins 8-13(-17) pairs, conspicuous abaxially, margin not revolute; perianth lobes densely pubescent inside.   14 P. neuranthoides
       
15 (1) Midrib of leaf blade completely elevated adaxially   (16)
+ Midrib of leaf blade completely or partly impressed adaxially   (18)
       
16 (15) Panicles usually 9-16 cm; leaf blade 9-22 cm, base attenuate and decurrent, lateral veins 10-15 pairs.   15 P. tavoyana
+ Panicles 3.5-10 cm; leaf blade 3-12 cm, base obtuse, not decurrent, lateral veins 6-8(-10) pairs   (17)
       
17 (16) Leaf blade lanceolate or oblanceolate, 8-12 × 3-3.5(-4) cm, veinlets conspicuous abaxially; inflorescences 6-10 cm.   16 P. legendrei
+ Leaf blade oblong or elliptic-oblong, 3-6 × 1.5-2 cm, veinlets slightly conspicuous abaxially; inflorescences ca. 3.5 cm.   17 P. brachythyrsa
       
18 (15) Midrib of leaf blade impressed at lower part only adaxially   (19)
+ Midrib of leaf blade completely impressed adaxially (but elevated distally in P. neurantha var. cavaleriei)   (20)
       
19 (18) Leaf blade obovate or oblanceolate, base attenuate and decurrent.   18 P. crassipedicella
+ Leaf blade lanceolate or elliptic-lanceolate, base cuneate, not decurrent.   16 P. legendrei
       
20 (18) Fruit globose.   19 P. forrestii
+ Fruit shape not as above   (21)
       
21 (20) Fruit large, more than 1.8 cm   (22)
+ Fruit smaller, below 1.5 cm   (27)
       
22 (21) One-year-old branchlets glabrous or puberulent   (23)
+ One-year-old branchlets conspicuously hairy   (24)
       
23 (22) Apex of leaf blade shortly acuminate, abaxially puberulent; fruit oblong or ellipsoid, 2-3.2 × 1.1-2 cm.   20 P. rufescens
+ Apex of leaf blade orbicular or slightly mucronate, abaxially usually glaucous; fruit long ovoid, ca. 1.8 × 1 cm.   21 P. glaucophylla
       
24 (22) Branchlets sparsely pubescent; leaf blade 10-20 cm, lateral veins 10-12 pairs.   22 P. yunnanensis
+ Branchlets densely yellowish brown tomentose or pubescent; leaf blade 15-45 cm, lateral veins more than 20 pairs   (25)
       
25 (24) Base of leaf blade shallowly cordate, oblique; fruit obovoid-oblong or obovoid; persistent perianth lobes ligneous, ca. 15 × 5-6 mm.   23 P. megacalyx
+ Base of leaf blade symmetrical; fruit ellipsoid; persistent perianth lobes leathery, 5-6 × 3.5-4 mm   (26)
       
26 (25) Leaf blade acuminate or shortly acuminate at apex, abaxially sparsely yellowish brown pubescent; fruit 3.5-3.8(-4.2) × 1.9-2.2 cm, apex glabrous.   24 P. macrocarpa
+ Leaf blade obtuse or mucronate at apex, abaxially densely yellowish brown villous especially along veins; fruit 2-2.2 × 1.5-1.6 cm, apex hairy.   25 P. motuonan
       
27 (21) One-year-old branchlets very thick, 4.5-6 mm in diam   (28)
+ One-year-old branchlets slender or very slender, 2-3.5 mm in diam   (29)
       
28 (27) Leaf blade obovate-lanceolate, elliptic-oblanceolate, or oblanceolate, 10-15 × 2-4.5 cm, lateral veins 12-14 pairs; branchlets, young leaf blade, petioles, and buds densely reddish brown or ferruginous villous.   26 P. hungmoensis
+ Leaf blade obovate-elliptic or broadly obovate-lanceolate, 10-23 × 5-9 cm, lateral veins 12-20 pairs; branchlets and petioles densely yellowish brown tomentose.   27 P. puwenensis
       
29 (27) Leaf blade lateral veins very slender, conspicuous or inconspicuous, veinlets nearly disappearing or completely disappearing abaxially; leaf blade gray-white appressed pubescent abaxially.   28 P. hui
+ Leaf blade lateral veins thick and veinlets conspicuous or very conspicuous abaxially; leaf blade hairy abaxially but not as above   (30)
       
30 (29) Fruit larger, 1.1-1.5 cm, ellipsoid-ovoid, ellipsoid, or suboblong; persistent perianth lobes clasping base of fruit   (31)
+ Fruit smaller, not more than 1 cm, ovoid or ellipsoid-ovoid; persistent perianth lobes loose, rarely clasping base of fruit   (33)
       
31 (30) Leaf blade obovate-elliptic or obovate-lanceolate, 7-17 × 3-7 cm; seed with numerous embryos, cotyledons unequal.   29 P. chekiangensis
+ Leaf blade elliptic, lanceolate, or oblanceolate, smaller; seed with solitary embryo, cotyledons equal   (32)
       
32 (31) Leaf blade leathery, lanceolate or oblanceolate, veinlets very distinct, reticulate abaxially; branchlets hairy or glabrate; panicles shorter, 3-7(-10) cm, not patent.   30 P. bournei
+ Leaf blade thinly leathery, elliptic, veinlets indistinct, not reticulate abaxially; branchlets densely yellowish brown or gray-brown pubescent; panicles longer, 7.5-12 cm, patent.   31 P. zhennan
       
33 (30) Older leaf blade abaxially and fruiting pedicel glabrous or sparsely pubescent   (34)
+ Older leaf blade abaxially and fruiting pedicel densely villous or pubescent   (35)
       
34 (33) Flowers smaller, 2-2.5 mm; persistent perianth lobes ca. 3 mm; leaf blade obovate or obovate-lanceolate, 9-15(-20) × 4-6(-8) cm, glaucous abaxially.   32 P. formosana
+ Flowers larger, 4-5 mm; persistent perianth lobes 4-5 mm; leaf blade lanceolate or oblanceolate, 3-16 × 1-4 cm.   33 P. neurantha
       
35 (33) Leaf blade densely or sparsely pubescent, ± glaucous abaxially; persistent perianth lobes unequal, clasping base of fruit.   34 P. glaucifolia
+ Leaf blade densely or sparsely gray-brown villous, not glaucous abaxially; persistent perianth lobes subequal, loose, not clasping base of fruit.   35 P. sheareri

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