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FOC | Family List | FOC Vol. 11 | Euphorbiaceae

9. Phyllanthodendron Hemsley, Hooker’s Icon. Pl. 26: ad t. 2563, 2564. 1898.

珠子木属 zhu zi mu shu

Authors: Bingtao Li & Michael G. Gilbert

Trees or shrubs, monoecious or dioecious, hairs simple, often absent. Leaves alternate, usually distichous; stipules small, caducous; petiole short; leaf blade simple, margin entire, venation pinnate. Inflorescences axillary, 1-flowered; pedicels often slender. Male flowers: sepals (4 or)5 or 6, free, in 2 series, imbricate, oblong-elliptic, midrib elevated abaxially, apex caudate-acuminate; petals absent; disk segments (4 or)5 or 6, shorter than sepals, ligulate or oblong, entire; stamens 3(or 4); filaments connate into a terete column; anthers 2-locular, longitudinally dehiscent, connectives subulate-acuminate at apex. Female flowers: sepals and disk segments 5 or 6; ovary 3-locular; ovules 2 per locule; styles 3, apex usually not lobed, often erect. Fruit a capsule, subglobose, septicidally and loculicidally dehiscent into 3 2-valved cocci when mature. Seeds 3-angled.

About 16 species: Peninsular Malaysia to China; ten species (nine endemic) in China.

Phyllanthodendron is a very easily identifiable segregate that nests within Phyllanthus, retained here for convenience. It can be easily distinguished by the distinctly caudate-acuminate sepals, linear to spatulate disk glands, and extended anther connectives.

1 Branches terete   (2)
+ Branches angular to prominently winged   (5)
2 (1) Branches, ovaries, and fruits pubescent or hispidulous; leaves abaxially brown or brown-red when dry   (3)
+ Branches, ovaries, and fruits glabrous; leaves abaxially glaucous when dry   (4)
3 (2) Young leaves and stipules abaxially puberulent, glabrescent; midrib and lateral veins raised on both surfaces.   1 P. yunnanense
+ Leaves and stipules abaxially glabrous; midrib and lateral veins adaxially flattened.   2 P. roseum
4 (2) Leaf blade obovate, papery, base cuneate or broadly so; lateral and reticulate veins abaxially flattened; disk segments spatulate; ovary globose, styles short, bifid to middle.   3 P. petraeum
+ Leaf blade rounded, leathery, base rounded or shallowly cordate; lateral and reticulate veins abaxially raised; disk segments ligulate or oblong; ovary ovoid, styles longer, bifid to nearly base.   4 P. orbicularifolium
5 (1) Branchlets angular, pubescent   (6)
+ Branchlets prominently winged, glabrous   (8)
6 (5) Leaf blade base oblique; male sepals 6.   5 P. breynioides
+ Leaf blade base symmetrical; male sepals 5   (7)
7 (6) Pedicels 1-2 mm; branches usually apically clustered; leaves of fertile branches 1-3 × 0.5-1.5 cm, leaf blade abaxially glaucous, lateral veins 6-8 pairs; veins of leaves, petioles, and stipules pilose; fruits glabrous.   6 P. anthopotamicum
+ Pedicels 10-25 mm; branches not clustered; leaves of fertile branches 6-12 × 2.5-4.5 cm, leaf blade abaxially greenish, lateral veins 8-12 pairs; veins of leaves, petioles, and stipules glabrous; fruits pubescent.   7 P. lativenium
8 (5) Male sepals, disk segments, and stamens 4; ovaries and fruits pubescent.   8 P. moi
+ Male sepals and disk segments 5 or 6, stamens 3; ovaries and fruits glabrous   (9)
9 (8) Leaf blade apex caudate, acumen ca. 1.2 cm, base cuneate or obtuse, lateral veins 10-14 pairs; male sepals and disk segments 6.   9 P. caudatifolium
+ Leaf blade apex acute or acuminate, base rounded, lateral veins 6-8 pairs; male sepals and disk segments 5.   10 P. dunnianum

Lower Taxa


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