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18. Physospermopsis H. Wolff, Notizbl. Bot. Gart. Berlin-Dahlem. 9: 276. 1925.

滇芎属 dian xiong shu

Pan Zehui (潘泽惠); Mark F. Watson

Haploseseli H. Wolff & Handel-Mazzetti.

Herbs perennial, glabrous. Taproot usually long, conic. Stem ribbed, base covered with fibrous sheath remnants. Leaf blade obovate-lanceolate, broadly triangular to oblong, 1–2-pinnate, rarely entire. Umbels terminal and lateral; bracts many, prominent, leaf-like, base entire, apex 3-lobed or pinnate; bracteoles present, entire or 3-lobed to pinnate. Calyx teeth minute or inconspicuous. Petals obovate, white, yellowish or dark purple, base shortly claw-like, apex obtuse-rounded or with shortly inflexed tips. Stylopodium flattened, margin sinuolate; styles about as long as stylopodium. Fruit ovoid to broadly ovoid, base slightly cordate, apex gradually narrowed, slightly flattened laterally, young fruits usually emerald green; ribs 5, filiform, prominent; vittae 2–3 in each furrow, 2–4 on commissure. Seed face plane to concave. Carpophore parted at apex.

About ten species: Sino-Himalayan region; eight species (four endemic) in China.

This is a taxonomically complex genus in which species boundaries are not always clear, and generic limits (e.g., with Pleurospermum, Tongoloa, and Trachydium) are problematic. Physospermopsis is one of a group of high-altitude Sino-Himalayan genera in need of a revision treating all taxa across their whole geographic range.


1 Basal leaves simple, undivided.   1 Physospermopsis alepidioides
+ Basal leaves 1–2-pinnate or 1–2-ternate/pinnate.   (2)
       
2 (1) Plants 5–10 cm high; stems reduced, often acaulescent.   2 Physospermopsis kingdon-wardii
+ Plants above 20 cm high; stems not reduced.   (3)
       
3 (2) Bracts usually absent; bracteoles inconspicuous.   3 Physospermopsis cuneata
+ and bracteoles present, conspicuous.   (4)
       
4 (3) Base of leaf segments decurrent, rachis broadly winged.   4 Physospermopsis delavayi
+ Base of leaf segments not decurrent, rachis not broadly winged.   (5)
       
5 (4) Basal leaves pinnate, nerves partly purple-red.   5 Physospermopsis rubrinervis
+ Basal leaves 2-pinnate, nerves not purple-red.   (6)
       
6 (5) Bracts entire, 3–5-lobed or nearly-pinnate.   6 Physospermopsis muliensis
+ Bracts 2-pinnate, as the leaves.   (7)
       
7 (6) Leaf sheaths broad-membranous; bracteoles narrowly obovate, apex incised into 3 teeth or pinnatifid; fruit ribs plane.   7 Physospermopsis shaniana
+ Leaf sheaths small, not broad-membranous; bracteoles broadly obovate, apex incised into 7–9 teeth; fruit ribs sinuolate.   8 Physospermopsis obtusiuscula

Lower Taxa


 

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